Leptosia

Leptosia Hübner, 1818

Leptosia Hübner, 1818; Zuträge Samml. exot. Schmett. 1: 13; TS: Leptosia chlorographa Hübner
Nina Horsfield, [1829]; Descr. Cat. lep. Ins. Mus. East India Coy (2): 140; TS: Papilio nina Fabricius
Nychitona Butler, 1870; Cistula ent. 1 (3): 34, 41; TS: Papilio dorothea Fabricius

Leptosia nina asukae; [iNaturalist]
Leptosia; Winhard, 2000, Butterflies of the world 10: 31; [CoL] ; [iNaturalist] ; [NHM-BMW] ; [NHM-BMW]
Leptosia (Anthocharidini); [AfrL]
Leptosia (Pierinae); Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 103
Nina; [NHM-BMW]
Nychitona; [NHM-BMW]
Nychitona (Pierinae); Moore, [1881], Lepid. Ceylon 1 (3): 117

Leptosia alcesta (Stoll, [1782])

Natal, Swaziland, Moçambique, Rhodesia, Tropical Africa. See [maps] Flip Flop African Wood White

Papilio alcesta Stoll, [1782]; in Cramer, Uitl. Kapellen 4 (32-32): 175, pl. 379, f. A, B
Papilio dorothea Fabricius, 1793; Ent. Syst. 3 (1): 194
?uganda Neustetter, 1927²; Int. ent. Z. 21: 7

Leptosia alcesta; [EOB]; [PBSA]; [EBW]; [AfrL]; [CoL] ; [iNaturalist] ; [LEPIFORUM]
?Leptosia uganda; [EBW]

L. a. alcesta W.Africa

Leptosia alcesta alcesta; [BAFR], 103 (text); [AfrL]; [iNaturalist]

L. a. inalcesta Bernardi, 1959 Ethiopia, Sudan, Uganda, Zaire, Kenya, Tanzania - Natal

Leptosia alcesta inalcesta; [BAFR], 103; [BK], 147; Winhard, 2000, Butterflies of the world 10: 31, pl. 48, f. 14; [AfrL]; [iNaturalist]

L. a. sylvicola (Boisduval, 1833) Madagascar

Leucophasia sylvicola Boisduval, 1833; ; TL: Madagascar

Leptosia alcesta sylvicola; [BAFR], 103 (text); [MadL]; [AfrL]; [iNaturalist]

L. a. pseudonuptilla Bernardi, 1959 Ethiopia

Leptosia alcesta pseudonuptilla; [AfrL]; [iNaturalist]
Leptosia nupta pseudonuptilla; [BAFR], 103 (text)
Leptosia pseudonuptilla; Winhard, 2000, Butterflies of the world 10: 31, pl. 48, f. 14

Leptosia bastini Hecq, 1997

Leptosia bastini; [AfrL]; [iNaturalist]

Leptosia hybrida Bernardi, 1952

-. See [maps] Hybrid Wood White

Leptosia hybrida; [AfrL]; [CoL] ; [iNaturalist]

L. h. hybrida Guinea - Zaire

Leptosia hybrida hybrida; [BAFR], 103; [AfrL]; [iNaturalist]

L. h. somereni Bernardi, 1959 E.Zaire, Uganda, W.Kenya

Leptosia hybrida somereni; [BAFR], 103 (text); [BK], 147; Winhard, 2000, Butterflies of the world 10: 31, pl. 48, f. 17; [AfrL]; [iNaturalist]

Leptosia marginea (Mabille, 1890)

Ivory Coast - Cameroun, W.Zaire. See [maps]

Leptosia marginea; [BAFR], 103; Winhard, 2000, Butterflies of the world 10: 31, pl. 48, f. 16; [AfrL]; [CoL] ; [iNaturalist]

Leptosia medusa (Cramer, [1777])

W.Africa, Sierra Leone - Ghana. See [maps]

Papilio medusa Cramer, [1777]; Uitl. Kapellen 2 (9-16): 86, pl. 150, f. F

Leptosia medusa; [EBW]; [BAFR], 103; [AfrL]; [CoL] ; [iNaturalist]
Pontia medusa; Boisduval, 1836, Hist. nat. Ins., Spec. gén. Lépid. 1: 433

Leptosia nina (Fabricius, 1793)

India, Ceylon, Burma, AU. See [maps] Psyche

Papilio nina Fabricius, 1793; Ent. Syst. 3 (1): 194, no. 604; TL: India
Papilio xiphia Fabricius, 1781; Spec. Ins. 2: 43 (preocc.)
Pontia crokera MacLeay, [1826]; in King, Narr. Surv. Austr. 2: 458
Leptosia xiphia micropunctata van Eecke, 1918; Zool. Meded. 4 (7): 73; TL: Pulu Lasia

Leptosia nina; [BIR]; [EBW]; [BOR], 122; [MRS], 262; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104; [CoL] ; [iNaturalist] ; [LEPIFORUM]
Leptosia xiphia; [BOW]: pl. 160, f. 29
Nychitona xiphia; Moore, [1881], Lepid. Ceylon 1 (3): 118
Nychitonia [= Nychitona] xiphia; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 836
Pontia crokera; Boisduval, 1836, Hist. nat. Ins., Spec. gén. Lépid. 1: 431
Pontia xiphia; Piepers & Snellen, 1909, Rhop. Java [1]: 3

769x513 (~65Kb) underside Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko

L. n. nina India, Ceylon - Indo-China, Thailand, Langkawi Is, Andamans

Leptosia nina nina; [BOR], 122; [BMP]: 85, pl. 8, f. 1; Winhard, 2000, Butterflies of the world 10: 31, pl. 48, f. 12; [Yutaka Inayoshi]

930x852 (~74Kb) Thailand, Chon Buri Province, Pattaya, a small grassy swamp with cattail and reed, surrounded by trees, at the Jomtien beach, between B.O. Guesthouse and Jomtien Metro Condotel. 26th January 2005, Photo © Oleg Kosterin	895x1156 (~207Kb) ♀ underside Cambodia, Ratanakiri Province, Lake Yak Lom, 30th May 2013, Photo © Oleg Kosterin	894x1158 (~237Kb) ♀ upperside Cambodia, Ratanakiri Province, Lake Yak Lom, 30th May 2013, Photo © Oleg Kosterin
1284x1375 (~266Kb) ♀ underside Cambodia, Ratanakiri Province, Lake Yak Lom, 30th May 2013, Photo © Oleg Kosterin

L. n. chlorographa Hübner, 1818 Java

Leptosia chlorographa Hübner, 1818; Zuträge Samml. exot. Schmett. 1: 13, pl. [9], f. 47-48; TL: Java

Leptosia nina chlorographa; [BOR], 122; [iNaturalist]

L. n. niobe (Wallace, 1866) Taiwan

Pontia niobe Wallace, 1866; Proc. zool. Soc. Lond. 1866 (2): 357; TL: Formosa

Leptosia nina niobe; [MRS], 262; [BOR], 122; [iNaturalist]

L. n. dione (Wallace, 1867) S.Sulawesi

Pontia dione Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3): 319; TL: Macassar

Leptosia nina dione; [BOR], 122; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104

L. n. nicobarica (Doherty, 1886) Nicobars

Nychitona xiphia var. nicobarica Doherty, 1886; J. asiat. Soc. Bengal 55 Pt.II (3): 262; TL: Nicobars

Leptosia nina nicobarica; [BOR], 122

L. n. fumigata Fruhstorfer, 1903 Lombok, Sumbawa, Flores, Solor

Leptosia xiphia fumigata Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2): 269; TL: Lombok
Leptosia xiphia fumigata Fruhstorfer, 1903; Soc. Ent. 18 (3): 18; TL: Lombok
?aurisparsa Fruhstorfer

Leptosia nina fumigata; [BOR], 122

L. n. terentia Fruhstorfer² S.Philippines

Leptosia nina terentia; [BOR], 122; [iNaturalist]
Leptosia terentia Fruhstorfer;

L. n. georgi Fruhstorfer² N.Philippines

Leptosia georgi Fruhstorfer;
Leptosia nina georgi; [BOR], 122; [iNaturalist]

L. n. comma Fruhstorfer, 1903 Timor - Tanimbar

Leptosia xiphia comma Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2): 269; TL: Timor; Wetter; Kalao; Tanah; Djampea; Selaru; Key
Leptosia xiphia comma Fruhstorfer, 1903; Soc. Ent. 18 (3): 18; TL: Timor; Wetter; Kalao; Tanah; Djampea; Selaru; Key

Leptosia nina comma; [BAUR], 124

L. n. malayana Fruhstorfer, 1910 Peninsular Malaya, Singapore, Sumatra, Borneo, Bangka, Biliton

Leptosia nina malayana Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 121, (44) ; TL: Malaysia

Leptosia nina malayana; [BOR], 122; [BMP]; [iNaturalist] ; [Yutaka Inayoshi]

L. n. aebutia Fruhstorfer, 1910² Tanahdjampea, Kalao

Leptosia nina aebutia; [BOR], 122; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104

Leptosia lignea (Vollenhoven, 1865)

Sulawesi. See [maps]

Pontia lignea Vollenhoven, 1865; Faune ent. 2 (Monogr. Pierid.): 4, pl. 2, f. 1a-b; TL: Celebes

Leptosia lignea; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104; [CoL] ; [iNaturalist]
Leptosia nina lignea; [BOR], 122; Winhard, 2000, Butterflies of the world 10: 31, pl. 48, f. 13

Leptosia nupta (Butler, 1873)

-. See [maps] Immaculate Wood White

Nychitona nupta Butler, 1873; Cist. Ent. 1 (7): 175; TL: Angola, Bembe Mines

Leptosia nupta; [AfrL]; [BK]; [CoL] ; [iNaturalist]

L. n. nupta Nigeria - Angola

Leptosia nupta nupta; [BK], 147; [BAFR], 103; [AfrL]; [iNaturalist]

L. n. pseudonupta Bernardi, 1959 Zaire, Uganda - Tanzania

Leptosia nupta pseudonupta; [BAFR], 103 (text); [AfrL]; [iNaturalist]

L. n. viettei Bernardi, 1959 Madagascar

Leptosia nupta viettei Bernardi, 1959; ; TL: Madagascar, Massif de l'Ankaratra, Forêt d'Ambahona, Manjakatompo

Leptosia nupta viettei; [BAFR], 103 (text); [MadL]; [AfrL]; [iNaturalist]

Leptosia wigginsi (Dixey, 1915)

-. See [maps] Opaque Wood White

Nychitona wigginsi Dixey, 1915; Trans. Ent. Soc. 1915 (1): 7, pl. 1, f. 9-12

Leptosia wigginsi; [BK], 148; [AfrL]; [CoL] ; [iNaturalist]

L. w. wigginsi Uganda, SW.Kenya, Tanzania, E.Zaire

Leptosia wigginsi wigginsi; [BAFR], 103; [AfrL]; [iNaturalist]

L. w. pseudalcesta Bernardi, 1965 Senegal - Ghana

Leptosia wigginsi pseudalcesta; [BAFR], 103 (text)#

Leptosia wigginsi pseudalcesta; [AfrL]; [iNaturalist]

References:

[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[NBC] Netherlands Centre for Biodiversity Naturalis
NBC naturalis
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Olli Vesikko]
http://koti.welho.com/ovesikko/
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[²] This may require parentheses or not (original genus not verified)

Some related literature:

Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, 1870
A revision of the Genera of the Sub-family Pierinae Cistula ent. 1 (3) : 33-58, pl. 1-4
Butler, 1873
Descriptions of new species of Lepidoptera Cist. Ent. 1 (7) : 151-177
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
[BMP]; Corbet & Pendlebury, 1992
The butterflies of the Malay Peninsula. Fourth Edition revised by J. N. Eliot with plates by Bernard D'Abrera Butterflies Malay Peninsula (4th edn) : 1-595, pl. 1-69
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
[BAUR]; D'Abrera, 1977
Butterflies of the Australian Region (2nd edn) Butts. Aust. Region: 1-416
[BAFR]; D'Abrera, 1980
Butterflies of the Afrotropical Region: -
[BOR]; D'Abrera, 1982-1986
Butterflies of the Oriental Region I-III
[PBSA]; Dickson & Kroon, 1978
Pennington's Butterflies of Southern Africa
Dixey, 1915
New species and subspecies of Pierinae Trans. Ent. Soc. 1915 (1) : 1-15, pl. 1-2
Doherty, 1886
Additional Notes on new or rare Indian Butterflies J. asiat. Soc. Bengal 55 Pt.II (3) : 256-265
van Eecke, 1918
Studies on Indo-Australian Lepidoptera--III. Some Rhopalocera and Netrocera[sic] from Simalur, Pulu Lasia, Pulu Babi and Sumatra Zool. Meded. 4 (7) : 70-101, 2pls
Fabricius, 1781
Species Insectorum Exhibentes Eorum Differentia Specifica, Synonyma Auctorum, Loca Natalia, Metamorphosin Adiectis, Observationibus, Descriptionibus. Tom II Spec. Ins. 2 : 1-494 (1781), : 495-514 (appendix), 515-517 (index) ([1782])
Fabricius, 1793
Entomologia Systematica emendata et aucta. Ent. Syst. 3 (1) : 1-487
[EOB]; Feltwell, 1993
The Encyclopedia of Butterflies
Fruhstorfer, 1903
Verzeichniss der in Tonkin, Annam und Siam gesammelten Pieriden und Besprechung verwandter Formen Dt. ent. Z. Iris 15 (2) : 269-305 (1 May)
Fruhstorfer, 1903
Verzeichniss der in Hinter-Indien gesammelten Pieriden Soc. Ent. 18 (3): 18-19 (1 May), (4): 25-26 (15 May)
Fruhstorfer, 1910
2. Familie: Pieridae, Weisslinge in Seitz, Gross-Schmett. Erde 9 : 119-120 (1909), : 121-144 (14 January, 1910), : 145-152 (11 February, 1910), : 153-168 (2 April, 1910), : 169-176 (4 April, 1910), : 177-190 (20 May, 1910)
Horsfield, [1829]
Descriptive Catalogue of the Lepidopterous Insects contained in the Museum of the Horourable East-India Company, illustrated by coloured figures of new species Descr. Cat. lep. Ins. Mus. East India Coy (1): 1-80, pl. 1-4 (1828), (2): 81-144, pl. 5-8, (expl.) [1-4] (1829)
Hübner, 1818
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 1 [1808-] 1818 Zuträge Samml. exot. Schmett. 1 : [1-3], 4-6, [7], 8-32, [33-40] (post 22nd December 1818), : pl. [1-2], f. 1-12 (1808 - 1809), : pl. [3-25], f. 13-146 (1809 - 1813), : pl. [26-35], f. 147-200 (1814 - 1818)
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
[MadL]; Lees, Kremen & Raharitsimba, 2003
Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist in Goodman & Benstead, Nat. Hist. Madagascar : 762-793
[BOW]; Lewis, 1974
Butterflies of the World
MacLeay, [1826]
in King, Narrative of a Survey of the Intertropical and Western Coasts of Australia. Catalogue of Insects, collected by Captain King, R.N. in King, Narr. Surv. Austr. 2 : 438-469
Moore, 1878
A list of the Lepidopterous Insects collected by Mr. Ossian Limborg in Upper Tenasserim, with Descriptions of new Species Proc. zool. Soc. Lond. 1878 (4) : 821-859, pl. 51-53 (1879)
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Neustetter, 1927
Neue afrikanische Tagfalter Int. ent. Z. 21 : 7-8, : 14-16, : 23-24, : 32
Piepers & Snellen, 1909
The Rhopalocera of Java. Pieridae Rhop. Java [1] : 1-65, pl. 1-4
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Stoll, [1782]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Vollenhoven, 1865
Essai d'une Faune Entomologique de l'Archipel Indo-Néerlandais, Seconde monographie: Famille des Piérides Faune ent. 2 (Monogr. Pierid.) : 1-70, pl. 1-7
Wallace, 1866
List of Lepidopterous insects collected at Takow, Formosa, by Mr. Robert Swinhoe Proc. zool. Soc. Lond. 1866 (2) : 355-365
Wallace, 1867
On the Pieridae of the Indian and Australian Regions Trans. ent. Soc. Lond. (3) 4 (3) : 301-416, pl. 6-9
Winhard, 2000
Pieridae I Butterflies of the world 10 : 1-40, pl. title,1-48,back
[BIR]; Wynter-Blyth, 1957
Butterflies of the Indian Region (1982 Reprint)

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