Coenonympha

Coenonympha Hübner, [1819]

Coenonympha Hübner, [1819]; Verz. bek. Schmett. (5): 65; TS: Papilio geticus Esper
Chortobius Dunning & Pickard, 1858; Accent. List Brit. Lep.: 5; TS: Papilio pamphilus Linnaeus
- = Coenonympha; [NL4A], 223; Pelham, 2008, J. Res. Lepid. 40: 398; [Wahlberg, 2023]
Chortobius Doubleday, 1859; Zool. Syn. List Br. Butts Moths (edn 2): 2 (preocc. Dunning & Pickard, [1859])
- = Coenonympha; [NL4A], 223; Pelham, 2008, J. Res. Lepid. 40: 398
Sicca Verity, 1953; Le Farfalle diurn. d'Italia 5: 83; TS: Papilio dorus Esper
- = Coenonympha; [NL4A], 223; Pelham, 2008, J. Res. Lepid. 40: 398; [Wahlberg, 2023]

Chortobius; [NHM-BMW] ; [NHM-BMW]
Coenonympha; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Coenonympha (Coenonymphina); [NL4A], 223; Pelham, 2008, J. Res. Lepid. 40: 398; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; [Wahlberg, 2023]
Coenonympha (Satyridae); Seitz, 1908, Gross-Schmett. Erde 1: 142; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 301
Coenonympha (Ypthimini); [RFEL]
Sicca; [NHM-BMW]

Coenonympha saadi Kollar, [1849]

Iraq, Iran, Kurdistan. See [maps]

Coenonympha saadi Kollar, [1849]; Denkschr. Akad. Wiss. Wien. 1: 52; TL: Shiraz, S.Iran

Coenonympha saadi; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 e; [BOW]: pl. 200, f. 2; [BRU]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

1169x1014 (~440Kb) underside Iran, Tehran, Elborz Mts. S foot, Tochal Mt. S. foot, Velenjak River valley, 2nd June 2017, Photo © Oleg Kosterin	880x1297 (~284Kb) underside Iran, Ostan-e Markazi, Shahrestan-e Arak, Anjedan (Anjudan) village, mountain foot, 22nd May 2017, Photo © Oleg Kosterin	1075x932 (~390Kb) underside Iran, Ostan-e Esfahan, the Ghomrood River left bank just upstream Golpayegan city, 21st May 2017, Photo © Oleg Kosterin
1199x1132 (~417Kb) underside Iran, Ostan-e Esfahan, the Ghomrood River left bank just upstream Golpayegan city, 21st May 2017, Photo © Oleg Kosterin	959x808 (~265Kb) underside Iran, Ostan-e Esfahan, the Ghomrood River left bank just upstream Golpayegan city, 21st May 2017, Photo © Oleg Kosterin

C. s. iphias Eversmann, 1851 Transcaucasia

Hipparchia (Coenonympha) iphias Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2): 618
mesopotamica Heyne, [1894]; Palaearkt. Grossschm. 1: 617

Coenonympha mesopotamica; [LepIndex] ; [CoL]
Coenonympha saadi mesopotamica; Seitz, 1908, Gross-Schmett. Erde 1: 145

Coenonympha mangeri Bang-Haas, 1927

Afghanistan. See [maps]

mangeri Bang-Haas, 1927; Horae Macrolep. Palaearct. 1: 50

Coenonympha mangeri; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

Coenonympha haydenii (Edwards, 1872)

Montana, Colorado. See [maps] Yellowstone Ringlet Wyoming Ringlet Hayden's Ringlet

Erebia haydenii Edwards, 1872; in Hayden, Geol. Surv. Montana: 467

Coenonympha haydeni; [BOW]: pl. 18, f. 14; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 399; [LepIndex] ; [CoL]
Coenonympha haydenii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30; [NACL], #4581; [NAB], 677; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]

Coenonympha symphita Lederer, 1870

NE.Turkey, SW.Georgia. See [maps]

Coenonympha symphita Lederer, 1870; Ann. Soc. ent. Belg. 13: 44; TL: Between Akhaltsikhe and the Turkish border, Georgia
?tiphonides Staudinger
Coenonympha symphita karsiana Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 351; TL: Transcaucasia
ab. inocellata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 352

Coenonympha symphita; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 g; [BRU], 194; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [LepIndex] ; [CoL] ; [iNaturalist]
?Coenonympha symphita tiphonides; Seitz, 1908, Gross-Schmett. Erde 1: 146

Coenonympha fettigii Oberthür, 1874

Morocco, Algeria, Tunisia. See [maps]

Coenonympha fettigii Oberthür, 1874; Petites Nouv. Ent. 1 (103): 412; TL: Province of Oran [Telaghre] (Algeria)

Coenonympha dorus fettigii; [H&R], 217
Coenonympha fettigii; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 f; [LepIndex] ; [CoL] ; [MBE] ; [MBE]

C. f. fettigii

Coenonympha fettigii fettigii ab. infra-simplex Rothschild, 1917; Novit. Zool. 24 (1): 118; TL: Les Pins (Algeria)

Coenonympha fettigii fettigii; [BMAT]: 84, pl. 29, f. 9-11

C. f. nicholasi Rothschild, 1925 Algeria (Blida, Chrea, etc.), Tunisia (Aïn Draham, Tunis)

Coenonympha dorus nicholasi Rothschild, 1925; Novit. Zool. 32: 208; TL: Blida-les-Glaciéres (Algeria)
Coenonympha fettigii holli Oberthür, 1910; Ètud. Lépid. Comp. 4: 42, pl. 48, f. 396-397; TL: Blida-les-Glaciéres (Algeria)

Coenonympha fettigii nicholasi; [BMAT]: 85, pl. 29, f. 12-17

C. f. inframaculata Oberthür, 1922 W.Morocco (Rif, Midle Atlas, High Atlas)

Coenonympha fettigi var. inframaculata Oberthür, 1922; Étud. Lépid. Comp. 19 (1): 87; TL: Fort Toumliline (Morocco)

Coenonympha fettigii inframaculata; [BMAT]: 86, pl. 29, f. 18-31

600x435 (~33Kb) underside Morocco, Djebel Tisiréne, Rif, Photo © Michel Tarrier

Coenonympha darwiniana Staudinger, 1871

SEU. See [maps]

Coenonympha arcania v. darwiniana Staudinger, 1871; Cat. Lepid. eur. Faunegeb. (Edn. 2): 32; TL: Valais, Switzerland

Coenonympha arcania darwiniana; [H&R], 219
Coenonympha arcania ab. darwiniana; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 d
Coenonympha darwiniana; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE]

Coenonympha mongolica Alphéraky, 1881 Ili River Basin

Coenonympha mongolica Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 426, 17 (1-2): pl. 1, f. 26; TL: Near Kuldzha, China

Coenonympha mongolica; Seitz, 1908, Gross-Schmett. Erde 1: 147, pl. 48 h; [BRU], 197; [BOW]: pl. 200, f. 10; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [SZMN]

Coenonympha oedippus (Fabricius, 1787)

CEU, S.Siberia - Ussuri, China, Korea, Japan. See [maps] Moor-Wiesenvögelchen False Ringlet Lindos ojos Strzępotek edypus

Papilio oedippus Fabricius, 1787; Mantissa Insectorum 2: 31; TL: S.Russia
Papilio geticus Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 51, (5-6): pl. 102, f. 2
?miris Fabricius
?hungarica ?Rebel, 1900
?oedippus ab. hungarica ?Aigner
Coenonympha oedippus; [BRU], 196

Coenonympha oedippus aquitanica; [LepIndex]; [CoL]
Coenonympha oedippus arothius; [LepIndex]; [CoL] ; [iNaturalist]
Coenonympha oedippus herbuloti; [LepIndex]; [CoL]
Coenonympha oedippus hungarica; [H&R], 223
Coenonympha oedippus reluminans; [LepIndex]; [CoL]
Coenonympha oedippus rhenana; [LepIndex]; [CoL] ; [iNaturalist]
Coenonympha oedippus rhodanica; [LepIndex]; [CoL]
Coenonympha oedippus sebrica; [LepIndex]; [CoL]
Coenonympha oedippus senonica; [LepIndex]; [CoL]
Coenonympha oedippus steni; [LepIndex]; [CoL]
Coenonympha oedippus; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 a; [H&R], 223; [BOW]: pl. 6, f. 4; [MRS], 403; [Otakar Kudrna]; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [Mario Maier]
?Coenonympha oedippus ab. miris; Seitz, 1908, Gross-Schmett. Erde 1: 143

1200x1565 (~150Kb) underside Hungary: Ocsa, June 2005, Photo © Szabolcs Safian

C. o. oedippus

Coenonympha oedippus oedippus; [H&R], 223; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46

C. o. annulifer Butler, 1877 Japan

Coenonympha annulifer Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109): 91; TL: 370 miles from Tokei (Yedo)
Coenonympha oedippus annulifer ab. oiwakensis Sugitani, 1932; Zephyrus 4 (1): 8

Coenonympha oedippus annulifer; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 a; Kudrna, 1974, Atalanta 5: 100; [iNaturalist]

C. o. magna Heyne, [1895] Altai, Tuva, ...

Coenonympha oedippus magna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; [iNaturalist]

600x800 (~97Kb) ♂ ♀ underside A meadow steppe on the hill Malyi Batur on the left bank of the Onon River 7 km upstream of the village Nizhnii Tsasuchei, Onon District, Chita Province, SE Transbaikalia (Dahuria), Siberia, Russia. 11th July 1996, Photo © Oleg Kosterin

857x1133 (~217Kb) underside Russia, West Siberia, the Altai Mts., Altai Republic, Shebalino District, the Ulus-Cherga village environs, a herbaceous meadow on southern slope. 11th July 2007 (on Phlomis tuberosa), Photo © Oleg Kosterin

1177x1309 (~266Kb) underside Russia, West Siberia, the Altai Mts., Altai Republic, Shebalino District, the Ulus-Cherga village environs, a herbaceous meadow on southern slope. 11th July 2007, Photo © Oleg Kosterin

C. o. amurensis Rühl, 1895 Primorye, ...

oedippus amurensis Rühl, 1895; Palaearkt. Grossschm. 1: 607

Coenonympha oedippus amurensis; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 a; [MRS], 403; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; [iNaturalist]

C. o. taibaica Murayama

Coenonympha oedippus taibaica; [MRS], 403

C. o. magnocellata Krzywicki, 1967

Coenonympha oedippus magnocellata Krzywicki, 1967; Ann. zool. Warsz. 25 (1): (1-213)

Coenonympha oedippus magnocellata; [LepIndex]; [CoL]

Coenonympha nolckeni Erschoff, 1874

Ghissar, Alai, Tian-Shan, N.Pamirs, W.Pamirs. See [maps]

Coenonympha nolckeni Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5): 23, pl. 2, f. 17; TL: Mt. Naubid, Zeravshan Valley, Uzbekistan

Coenonympha nolckeni; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 b; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 497; [BRU], 196; [BOW]: pl. 200, f. 9; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; Tshikolovets, 2017, Zootaxa 4358 (1): 119; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

Coenonympha xinjiangensis Chou & Huang, 1994

Xinjiang. See [maps]

Coenonympha xinjiangensis Chou & Huang, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 759, 403, f. 32-33; TL: Xinjiang, Jiandengyu

Coenonympha tydeus Leech, [1892]

Tibet. See [maps]

Coenonympha tydeus Leech, [1892]; Butts China Japan Corea (1): 96

Coenonympha amaryllis tydeus; [iNaturalist]
Coenonympha tydeus; [MRS], 405; [iNaturalist]
?Coenonympha tyderes[sic, tydeus?]; [BOW]: pl. 200, f. 5 (text only)

Coenonympha sinica Alphéraky, 1888

Kashmir, Tibet. See [maps]

Coenonympha sinica Alphéraky, 1888; Stettin Ent. Ztg 49: 66; TL: Nian-Chian-Kette, Djin-ta-sy

Coenonympha sinica; Seitz, 1908, Gross-Schmett. Erde 1: 147, pl. 48 h; [MRS], 405; [LepIndex] ; [CoL] ; [iNaturalist]

Coenonympha pavonina Alphéraky, 1888

China. See [maps]

Coenonympha pavonina Alphéraky, 1888; Stettin Ent. Ztg 49: 66; TL: R. Hei-ho

Coenonympha pavonina; Seitz, 1908, Gross-Schmett. Erde 1: 146; [LepIndex] ; [CoL]

The tullia species-group

group Tullia (Coenonympha); Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318

Coenonympha dorus (Esper, 1782)

NAF, SWEU, it. See [maps] Dusky Heath Velago de negro

Papilio dorus Esper, 1782; Die Schmett. Th. I, Bd. 2 (5): 130, pl. 78 f. 1
Coenonympha dorus v. bieli Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 65; TL: Portugal
?andalusica Ribbe
?caeca Oberthür
?fulvia Oberthür
Coenonympha mathewi Tutt, 1904; Ent. Rec. J. Var. 16: 308; TL: Vigo
Coenonympha dorus fonti de Sagarra, 1924; Butll. Inst. Catal. Hist. nat. (2) 4 (9): 199; TL: Albarracin (Aragó), 1100m
dorus aquilonia Higgins, 1969; Entomologist 10 (1270): 60; TL: l'Aquila, Abruzzi, Italy

Coenonympha dorus andalusica; Seitz, 1908, Gross-Schmett. Erde 1: 145
Coenonympha dorus aquilonia; [H&R], 217; [LepIndex]; [CoL]
Coenonympha dorus bieli; Seitz, 1908, Gross-Schmett. Erde 1: 145; [H&R], 217
Coenonympha dorus cantabrica; [LepIndex]; [CoL]
Coenonympha dorus fulvia; Seitz, 1908, Gross-Schmett. Erde 1: 145
Coenonympha dorus macrophthalma; [LepIndex]; [CoL]
Coenonympha dorus purpurinatta; [LepIndex]; [CoL]
Coenonympha dorus; [H&R], 216; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 f; [EBW]; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BMAT]; [BOW]: pl. 5, f. 17; [Otakar Kudrna]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [Mario Maier]
Coenonympha dorus ab. caeca; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 f

568x740 (~90Kb) Albarracin, prov. Teruel, , Spain, August-1988, Photo © Enrique Garcia-Barros

C. d. dorus

Coenonympha dorus dorus; [H&R], 216

C. d. austauti Oberthür, 1881 W.Algeria Austaut's Algerian Heath

Coenonympha dorus var. austauti Oberthür, 1881; Étud. d'Ent. 6: 59; TL: Nemours [Ghazaouet] (Algeria)

Coenonympha austauti; [H&R], 218; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note)
Coenonympha dorus austauti; Seitz, 1908, Gross-Schmett. Erde 1: 145; [BMAT]: 86, pl. 29, f. 1-8

Coenonympha pamphilus (Linnaeus, 1758)

Palearctic. See [maps] Wiesenvögelchen Kamgräsfjäril Kälberauge Small Heath Strzępotek ruczajnik Staggängefjäril Nispola Le Procris Keltaniittyperhonen Kleiner Heufalter

Papilio pamphilus Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 472; TL: Sweden
?Coenonympha pamphiloides Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1): 146
- = ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30
?nephela Hufnagel
- = ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30
?menalcas Poda, 1761; Ins. Mus. Graecensis: 78
- = ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30
Coenonympha pamphilus orantia Fruhstorfer, 1908; Int. ent. Zs. 2 (2): 10; TL: Kashgar
?bipupillata Cosmovici
?Coenonympha pamphilus ab. thyrsides Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 66
?Coenonympha pamphilus ab. semilyllus Krulikowsky, 1908; Soc. Ent. 23 (1): 3; TL: Wiatka; Kasan
Coenonympha pamphilus ab. amaryllides Stauder, 1911; Boll. Soc. Adriat. 25: (?)
Coenonympha pamphilus australis Verity, 1914; Boll. Soc. ent. ital. 45: 227
Coenonympha pamphilus ab. bipunctata Gussich, 1917; Godina 29: 221; TL: Croatia
Coenonimpha[sic] pamphilus altopirenaica de Sagarra, 1930; Butll. Inst. catal. Hist. nat. (2) 10 (7): 113

Coenonympha pamphilius[sic, pamphilus]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30
Coenonympha pamphiloides; [LepIndex] ; [CoL]
Coenonympha pamphilus altopirenaica; [LepIndex]; [CoL]
Coenonympha pamphilus neolyllus; [LepIndex]; [CoL]
Coenonympha pamphilus wimbladi; [LepIndex]; [CoL]
Coenonympha pamphilus; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 497; Seitz, 1908, Gross-Schmett. Erde 1: 146; [SPRK]; [H&R], 214; [BRU], 192; [EBW]; [BOW]: pl. 6, f. 5; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [MRS], 404; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; Yakovlev, 2012, Nota lepid. 35 (1): 78; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF] ; [Mario Maier]
?Coenonympha pamphilus thyrsides; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 g
?Coenonympha pamphilus ab. bipupillata; Seitz, 1908, Gross-Schmett. Erde 1: 146

500x376 (~39Kb) underside Hoe lake, Devon, UK 24/06/95, Photo © Simon Coombes	1000x1076 (~169Kb) underside Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 8.8.2004, Photo © Markku Savela	700x572 (~77Kb) underside France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela
700x603 (~74Kb) underside France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela	470x384 (~99Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý	1200x900 (~187Kb) underside Italy, Ravenna, August 2002, Photo © Maurizio Elia
293x244 (~10Kb) ♀	282x259 (~10Kb) ♂	900x723 (~54Kb) underside FINLAND, Ab: Suomusjärvi, 669:31, 28.6.1998 Photo © Markku Savela

C. p. pamphilus

Coenonympha pamphilus pamphilus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 g

C. p. lyllus (Esper, 1806)

Papilio lyllus Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 23, pl. 122, f. 1-2; TL: Portugal
Coenonympha pamphilus lyllus gen. vern. lyllides Verity, 1914; Boll. Soc. ent. ital. 45: 226
Coenonympha pamphilus latevittata Verity, 1916; Ent. Rec. J. Var. 28 (7-8): 172; TL: Africa [Algeria]
Coenonympha pamphilus arenosa Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 198; TL: Fez (Morocco)
Coenonympha pamphilus atlantea Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 199; TL: Atlas Mts. (Morocco)

Coenonympha pamphilus lyllus; Seitz, 1908, Gross-Schmett. Erde 1: 146; [BMAT]: 84, pl. 28, f. 28-44; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; [iNaturalist]

600x435 (~30Kb) underside Morocco, Pays Beni-Routen, Rif, Photo © Michel Tarrier

C. p. marginata Heyne, [1894]

pamphilus marginata Heyne, [1894]; Palaearkt. Grossschm. 1: 619
?marginata Staudinger

Coenonympha pamphilus marginata; [iNaturalist]
?Coenonympha pamphilus marginata; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 g

C. p. ferghana Stauder, 1924

pamphilus ferghana Stauder, 1924; Int. ent. Zs. 17: 152

C. p. fulvolactea Verity, 1926

pamphilus fulvolactea Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 205

C. p. centralasiae Verity, 1926

pamphilus centralasiae Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 205

C. p. infrarasa Verity, 1926

pamphilus infrarasa Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 206

C. p. juldusica Verity, 1926

pamphilus juldusica Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 206

C. p. nitidissima Verity, 1926

pamphilus nitidissima Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 199

C. p. asiaemontium Verity, 1926

pamphilus asiaemontium Verity, 1926; Zs. Wiss. InsektBiol. Berlin 21: 206

Coenonympha thyrsis (Freyer, 1845) Crete

thyrsis Freyer, 1845; Neuere Beitr. Schmett. 5: pl. 475, f. 1

Coenonympha thyrsis; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 e; [Otakar Kudrna]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]

Coenonympha tullia (Müller, 1764)

NWEU, Temperate Asia, NA. See [maps] Saraikkoniittyperhonen Le Daphnis Large Heath Strzępotek soplaczek Starrgräsfjäril Starrängefjäril Common Ringlet Grosser Heufalter

Papilio tullia Müller, 1764; Fauna Ins. Fridr.: 36; TL: Zealand Is., Denmark
Papilio davus Fabricius, 1777; Genera Insectorum: 259
Papilio isis Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 31
- = ; Pelham, 2008, J. Res. Lepid. 40: 399
Coenonympha davus var. rothliebi Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 14
Coenonympha tiphon v. scotica Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 66; TL: Scotia
?occupata Rebel, 1903; Ann. nat. Mus. Wien 18: 182
Coenonympha typhon fermana Fruhstorfer, 1908; Int. ent. Zs. 2 (2): 10; TL: Kashgar
Coenonympha typhon thimoites Fruhstorfer, 1910; Soc. ent. 25 (14): 54; TL: Martigny
Coenonympha tullia muonioensis Lingonblad, 1950; Notulae Ent. 29 (3-4): 81; TL: Finland, Muonio

Coenonympha tiphon isis; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 h
Coenonympha tiphon scotica; Seitz, 1908, Gross-Schmett. Erde 1: 146
Coenonympha tullia davus; [iNaturalist]
Coenonympha tullia fiorii; [CoL]
Coenonympha tullia muonioensis; [CoL]
Coenonympha tullia muonioënsis; [LepIndex]
Coenonympha tullia rothliebi; [H&R], 213
Coenonympha tullia scotica; [H&R], 213
Coenonympha tullia; [BRU], 193; [H&R], 212; [EBW]; [BOW]: pl. 6, f. 6; [Opler]; [Otakar Kudrna]; Pelham, 2008, J. Res. Lepid. 40: 399; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; Yakovlev, 2012, Nota lepid. 35 (1): 78; [RFEL]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF] ; [Mario Maier]
?Coenonympha tiphon occupata; Seitz, 1908, Gross-Schmett. Erde 1: 146

599x400 (~31Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	400x599 (~27Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch	400x588 (~38Kb) underside Central Europe 1990-2000, Photo © Michael Borsch
444x358 (~40Kb) underside USA: Alabama, 2.8.1999, Photo © Vitaly Charny	351x621 (~35Kb)	360x285 (~19Kb) ♀
370x282 (~21Kb) ♂

C. t. tullia EU, W.Siberia

Coenonympha tiphon Rottemburg, 1775; Der Naturforscher 6: 15
- = Coenonympha tullia; Pelham, 2008, J. Res. Lepid. 40: 399
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47
Papilio philoxenus Esper, 1780; Die Schmett. Th. I, Bd. 2 (1): 25, pl. 54, f. 3
- = ; [H&R], 212
?Papilio laidion Borkhausen, 1788; Naturges. Eur. Schmett. 1: 91
Coenonympha tullia timanica Sedykh, 1977; Geogr. Aspect. Conserv. Flora Faun. NE Europ. USSR: 102
- = Coenonympha tullia; Pelham, 2008, J. Res. Lepid. 40: 399

Coenonympha tiphon philoxenus; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 h
Coenonympha tiphon; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 h
Coenonympha tullia tiphon; [H&R], 213
Coenonympha tullia tullia; [H&R], 212; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [iNaturalist]
?Coenonympha tiphon ab. laidion; Seitz, 1908, Gross-Schmett. Erde 1: 146
?Coenonympha typhon laidon[sic, laidion]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30
?Coenonympha typhon[sic, tiphon]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30

C. t. demophile (Freyer, 1845) NEU, N.Siberia (Yamal Peninsula)

Papilio (Hipparchia) demophile Freyer, 1845; Neuere Beitr. Schmett. 5: 97, pl. 439, f. 3-4
- = Coenonympha tullia; Pelham, 2008, J. Res. Lepid. 40: 399
Coenonympha tullia suevica Hemming, 1936; Proc. R. ent. Soc. Lond. (B) 5 (6): 123 (repl. Hipparchia demophile Freyer, 1845)
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47
- = Coenonympha tullia; Pelham, 2008, J. Res. Lepid. 40: 399
?demophile (Hübner, [1790-1793]) (Papilio); Schmett. Lep. linn.: 10 (ICZN 975)
fridolini Kusnezov, 1941; in Davenport
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47
minimus Sedykh, 1977

Coenonympha tullia demophile; [H&R], 214; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. t. viluiensis Ménétriés, 1859 Chukot Peninsula

witimensis Davenport, 1941; Bull. Mus. comp. Zool. Harv. 87 (4): 253
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

Coenonympha tiphon viluiensis; Seitz, 1908, Gross-Schmett. Erde 1: 146
Coenonympha tullia viluiensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [iNaturalist]
Coenonympha tullia witimensis; [LepIndex]; [CoL]

C. t. inornata Edwards, 1861 Quebec, New York Prairie Ringlet

Coenonympha inornata Edwards, 1861; Proc. Acad. nat. Sci. Philad. 13: 160, 163; TL: Lake Winnipeg
Coenonympha inornata r. quebecensis Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 89; TL: Chelsea, Ottawa Co., Quebec
- = ; Pelham, 2008, J. Res. Lepid. 40: 399
Coenonympha nipisiquit heinemani Brown, [1959]; J. N.Y. Ent. Soc. 66 (1/2): 67, pl. 2; TL: Grindstone I., Clayton, Jefferson Co., New York
- = ; Pelham, 2008, J. Res. Lepid. 40: 400

Coenonympha california inornata; [P&N23]; [iNaturalist]
Coenonympha inornata heinemani; [NACL], #4583d; [LepIndex]; [CoL]
Coenonympha inornata macisaaci; [LepIndex]; [CoL]
Coenonympha inornata; [NACL], #4583; [BOW]: pl. 18, f. 7; [LepIndex] ; [CoL]
Coenonympha tullia inornata; [USGS]; Pelham, 2008, J. Res. Lepid. 40: 399

C. t. ochracea Edwards, 1861 Colorado Ocher Ringlet

Coenonympha ochracea Edwards, 1861; Proc. Acad. nat. Sci. Philad. 13: 160, 163; TL: Turkey Creek, Jefferson Co., Colorado, 8000 (neotype)
Coenonympha ochracea tr. f. albescens Field, 1936; J. Ent. Zool. 28 (2): 22; TL: Albany Co., Wyoming
- = ; Pelham, 2008, J. Res. Lepid. 40: 402
Coenonympha tullia phantasma Burdick, 1956; Lepid. News 10 (3/4): 81; TL: Eldora, Boulder Co., Colorado
- = ; Pelham, 2008, J. Res. Lepid. 40: 402

Coenonympha california ochracea; [P&N23]; [iNaturalist]
Coenonympha inornata phantasma; [LepIndex]; [CoL]
Coenonympha ochracea; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30; [NACL], #4584; [NL4A], #1602; [LepIndex] ; [CoL]
Coenonympha tullia ochracea; [USGS]; Pelham, 2008, J. Res. Lepid. 40: 402
ochracia[sic, ochracea]; Chermock & Chermock, 1938

C. t. kodiak Edwards, 1869 Alaska, Yukon, NW.Territories, N.Siberia Kodiak Ringlet

Coenonympha kodiak Edwards, 1869; Trans. amer. ent. Soc. 2 (4): 375; TL: Kodiak, Alaska

Coenonympha kodiak; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30; [NACL], #4582; [LepIndex] ; [CoL]
Coenonympha tullia kodiak; Pelham, 2008, J. Res. Lepid. 40: 400; [P&N23]; [iNaturalist] ; [BOA]

C. t. ampelos Edwards, 1871 Northwest Ringlet

Coenonympha ampelos Edwards, 1871; Trans. amer. ent. Soc. 3 (3/4): 213
Coenonympa sweadneri Chermock & Chermock, 1941; Proc. Penn. Acad. Sci. 15: 42; TL: Pine Creek, Shoshone Co., Idaho
- = ; Pelham, 2008, J. Res. Lepid. 40: 401

Coenonympha ampelos; [NACL], #4585
Coenonympha california ampelos; [P&N23]; [iNaturalist]
Coenonympha sweadneri; [LepIndex] ; [CoL]
Coenonympha tullia ampelos; [USGS]; Pelham, 2008, J. Res. Lepid. 40: 401

identification uncertain 700x510 (~33Kb) underside USA: Lake Sammamish Park, Issaquah (47°33'36N 122°03'25W), King Co., WA, 11.7.1999, Photo © Markku Savela	identification uncertain 800x739 (~47Kb) underside USA: Issaquah (47°32'35N 122°04'00W), King Co., WA, 7.7.1999, Photo © Markku Savela	identification uncertain 500x439 (~34Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela
identification uncertain 900x1002 (~120Kb) underside USA: near Tyler (47°27'09"N 117°48'41"W), Spokane Co., Washington, 18.8.2005, Photo © Markku Savela

C. t. eryngii H. Edwards, 1877 California

Coenonympha californica[sic] var. eryngii H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 (12): 172; TL: Soda Springs, Siskiyou Co., California
Coenonympha california siskiyouensis Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 62; TL: Mt Shasta, California
- = ; Pelham, 2008, J. Res. Lepid. 40: 401

Coenonympha california eryngii; [NACL], #4586a; [P&N23]; [iNaturalist] ; [BOA]
Coenonympha tullia eryngii; Pelham, 2008, J. Res. Lepid. 40: 401

C. t. elko Edwards, 1881 Nevada - Vancouver

Coenonympha elko Edwards, 1881; Can. Ent. 13 (3): 57; TL: Elko, Nevada

Coenonympha ampelos elko; [NACL], #4585b
Coenonympha california elko; [P&N23]; [iNaturalist]
Coenonympha elko; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30; [BOW]: pl. 18, f. 6; [LepIndex] ; [CoL]
Coenonympha tullia elko; Pelham, 2008, J. Res. Lepid. 40: 401

C. t. caeca Staudinger, 1886 Pamirs-Alai, Ghissar, Tian-Shan - Altai

Coenonympha caeca Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 251; TL: Chatkalsky Mts, Uzbekistan

Coenonympha caeca; [BRU], 193; [LepIndex] ; [LEPIFORUM]
Coenonympha tiphon caeca; Seitz, 1908, Gross-Schmett. Erde 1: 146
Coenonympha tullia caeca; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; [iNaturalist]

C. t. subcaeca Heyne, [1894] Sailyughem Mts, Altai Mts.

tullia subcaeca Heyne, [1894]; Palaearkt. Grossschm. 1: 827
Coenonympha tullia elwesi Davenport, 1941; Bull. Mus. comp. Zool. Harv. 87 (4): 249; TL: Altai Mts
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

Coenonympha iphis subcaecata[sic, subcaeca]; Seitz, 1908, Gross-Schmett. Erde 1: 147
Coenonympha tullia elwesi; [LepIndex]; [CoL] ; [iNaturalist]
Coenonympha tullia subcaeca; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [iNaturalist]

elwesi
413x400 (~45Kb) ♂ underside An alpine meadow, at 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin	585x733 (~252Kb) ♂ underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian} Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 3rd July 2007, Photo © Oleg Kosterin

C. t. mixturata Alphéraky, 1897 Kamchatka

Coenonympha tiphon var. mixturata Alphéraky, 1897; in Romanoff, Mém. Lép. 9: 326, pl. 14, f. 4

Coenonympha kodiak mixturata; [NACL], #4582a
Coenonympha tiphon mixturata; Seitz, 1908, Gross-Schmett. Erde 1: 146
Coenonympha tullia mixturata; [P&N23]; [iNaturalist] ; [BOA]
Coenotympha tullia mixturata; [RFEL]; Pelham, 2008, J. Res. Lepid. 40: 400; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. t. yukonensis Holland, 1900 Alaska

Coenonympha kodiak var. yukonensis Holland, 1900; Ent. News 11 (3): 386; TL: Dawson, Yukon Territory ... Eagle City, Alaska .. American Creek

Coenonympha kodiak yukonensis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30; [NACL], #4582b
Coenonympha tullia yukonensis; Pelham, 2008, J. Res. Lepid. 40: 400; [P&N23]; [iNaturalist] ; [BOA]

C. t. eupompus Stauder, 1924 Dzhungarsky Alatau

pamphilus eupompus Stauder, 1924; Int. ent. Zs. 17: 152; TL: Ili
Coenonympha caeca heptopotamica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 352; TL: Aulie-ata

Coenonympha tullia eupompus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. t. furcae Barnes & Benjamin, 1926 Arizona

Coenonympha furcae Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 90; TL: Grand Canyon, Arizona

Coenonympha california furcae; [P&N23]; [iNaturalist]
Coenonympha ochracea furcae; [NACL], #4584d

C. t. subfusca Barnes & Benjamin, 1926 Arizona

Coenonympha ochracea r. subfusca Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 90; TL: White Mts, Arizona

Coenonympha california subfusca; [P&N23]; [iNaturalist]
Coenonympha ochracea subfusca; [NACL], #4584e
Coenonympha tullia subfusca; Pelham, 2008, J. Res. Lepid. 40: 402

C. t. benjamini McDunnough, 1928 Alberta

Coenonympha inornata benjamini McDunnough, 1928; Can. ent. 60 (11): 272; TL: Waterton Lakes, Alberta
Coenonympha tullia bottineauensis Chermock, Simmons & Chermock, 1963; North Dakota Insects 2: 23
- = ; Pelham, 2008, J. Res. Lepid. 40: 400

Coenonympha california benjamini; [P&N23]; [iNaturalist]
Coenonympha inornata benjamini; [NACL], #4583b
Coenonympha tullia benjamini; Pelham, 2008, J. Res. Lepid. 40: 400

C. t. columbiana McDunnough, 1928 British Columbia

Coenonympha inornata columbiana McDunnough, 1928; Can. ent. 60 (11): 273; TL: Aspen Grove, B.C.

Coenonympha ampelos columbiana; [NACL], #4585c
Coenonympha california columbiana; [P&N23]; [iNaturalist]
Coenonympha tullia columbiana; Pelham, 2008, J. Res. Lepid. 40: 401

C. t. insulanus McDunnough, 1928 British Columbia (Vancouver Is.)

Coenonympha inornata insulanus McDunnough, 1928; Can. ent. 60 (11): 273; TL: Victoria, B.C

Coenonympha ampelos insulana; [NACL], #4585d
Coenonympha california insulana; [P&N23]; [iNaturalist]
Coenonympha tullia insulanus; Pelham, 2008, J. Res. Lepid. 40: 400

C. t. mcisaaci dos Passos, 1935 Newfoundland

Coenonympha inornata mcisaaci dos Passos, 1935; Can. Ent. 67 (4): 83; TL: Doyles Station, Newfoundland
macisaaci [= mcisaaci] dos Passos, 1964; (emend.)
- = ; Pelham, 2008, J. Res. Lepid. 40: 399

Coenonympha california mcisaaci; [P&N23]; [iNaturalist]
Coenonympha inornata mcisaaci; [NACL], #4583a
Coenonympha tullia mcisaaci; Pelham, 2008, J. Res. Lepid. 40: 399

C. t. mackenziei Davenport, 1936

Coenonympha tiphon mackenziei Davenport, 1936; Can. Ent. 68 (4): 79

Coenonympha california mackenziei; [P&N23]; [iNaturalist]
Coenonympha ochracea mackenziei; [NACL], #4584b
Coenonympha tullia mackenziei; Pelham, 2008, J. Res. Lepid. 40: 402; [LepIndex]; [CoL]

C. t. chatiparae Sheljuzhko, 1937 Caucasus Minor

Coenonympha tullia chatiparae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 46; [LepIndex]; [CoL] ; [iNaturalist]

730x1025 (~173Kb) underside Russian Federation, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, the Blyam Mt. environs, 12 July 2008, Photo © Oleg Kosterin	662x882 (~150Kb) underside Russian Federation, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, the Blyam Mt. environs, 12 July 2008, Photo © Oleg Kosterin	959x1119 (~262Kb) underside Russian Federation, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, the Blyam Mt. environs, 12 July 2008, Photo © Oleg Kosterin
649x956 (~145Kb) underside Russian Federation, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, the Blyam Mt. environs, 12 July 2008, Photo © Oleg Kosterin

C. t. nipisiquit McDunnough, 1939 New Brunswick, Quebec Nipisquit Ringlet

Coenonympha inornata var. nipisiquit McDunnough, 1939; Can. ent. 71 (12): 266; TL: near Bathurst, New Brunswick

Coenonympha california nipisiquit; [P&N23]; [iNaturalist]
Coenonympha inornata nipisiquit; [NACL], #4583c
Coenonympha nipisiquit;
Coenonympha tullia nipisiquit; Webster, 1998, J. Lep. Soc. 52 (4): 345-355; Pelham, 2008, J. Res. Lepid. 40: 399

C. t. bosniae Davenport, 1941

Coenonympha tullia bosniae Davenport, 1941; Bull. Mus. comp. Zool. Harv. 87 (4): 244; TL: Lake Jesero, Bosnia

Coenonympha tullia bosniae; [LepIndex]; [CoL]
bosniae; Abadjiev, 2002, Neue Ent. Nachr. 53: 27

C. t. sibirica Davenport, 1941 Amur

Coenonympha tullia sibirica Davenport, 1941; Bull. Mus. comp. Zool. Harv. 87 (4): 254 (?preocc. Coenonympha hero var. sibirica Fuchs, 1899); TL: Seja, Amur region

Coenonympha tullia sibirica; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [LepIndex]; [CoL] ; [iNaturalist]

C. t. mono Burdick, 1942 California

Coenonympha ampelos r. mono Burdick, 1942; Can. Ent. 74 (1): 2; TL: Bridgeport, Mono Co., California

Coenonympha california mono; [P&N23]; [iNaturalist]
Coenonympha ochracea mono; [NACL], #4584c
Coenonympha tullia mono; Pelham, 2008, J. Res. Lepid. 40: 401

C. t. eunomia Dornfeld, 1967 Oregon, ...

Coenonympha tullia eunomia Dornfeld, 1967; J. Lepid. Soc. 21 (1): 3; TL: Wilhoit, Oregon
Coenonympha ampleos f. eunomia Field, 1937; Can. Ent. 69 (11): 249; TL: Wilhoit, Oregon
- = ; Pelham, 2008, J. Res. Lepid. 40: 401

Coenonympha ampelos eunomia; [NACL], #4585a
Coenonympha california eunomia; [P&N23]; [iNaturalist]
Coenonympha tullia eunomia; Pelham, 2008, J. Res. Lepid. 40: 401; [LepIndex]; [CoL]

C. t. yontocket Porter & Mattoon, 1989 N.California (Del Norte Co.)

Coenonympha tullia yontocket Porter & Mattoon, 1989; J. Lep. Soc. 43 (3): 231, f. 1-3; TL: California, Del Norte Co. (dunes N of Crescent City, between Lake Earl and Smith River)

Coenonympha california yontocket; [P&N23]; [iNaturalist]
Coenonympha tullia yontocket; Pelham, 2008, J. Res. Lepid. 40: 400

C. t. pseudobrenda Austin & Gray, 1998 Nevada

Coenonympha tullia pseudobrenda Austin & Gray, 1998; Syst. W. N. Am. Butts. (47): 588; TL: Nevada, Lincoln Co., Wilson Creek Range, road towards Wilson Peak from Atlanta Road, 9.6 road miles south of Atlanta Road, 2470m, T5N R68E S22 on USGS Trail Canyon 15' quadrangle

Coenonympha california pseudobrenda; [P&N23]; [iNaturalist]
Coenonympha tullia pseudobrenda; Pelham, 2008, J. Res. Lepid. 40: 401

C. t. tshonkurtshakus Korb, 1999

Coenonympha tullia tshonkurtshakus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. t. streltzovi Dubatolov, 2005

Coenonympha tullia streltzovi Dubatolov, 2005; 251

Coenonympha california Westwood, [1851]

California. See [maps] California Ringlet

Coenonympha california Westwood, [1851]; Gen. diurn. Lep. (2): 398; TL: California
Satyrus galactinus Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 309; TL: California
- = Coenonympha california california; [NL4A], #1601a
- = Coenonympha tullia california; Pelham, 2008, J. Res. Lepid. 40: 400
Coenonympha ceres Butler, 1866; Ent. mon. Mag. 3 (28): 78; TL: California
- = Coenonympha california california; [NL4A], #1601a
- = Coenonympha tullia california; Pelham, 2008, J. Res. Lepid. 40: 400
Coenonympha californica[sic] var. pulla H. Edwards, 1881; Papilio 1 (4): 51; TL: San Mateo Co., California
- = Coenonympha california california; [NL4A], #1601a
- = Coenonympha tullia california; Pelham, 2008, J. Res. Lepid. 40: 401
Caenonympha[sic] brenda Edwards, 1869; Trans. amer. ent. Soc. 2 (4): 375; TL: California
- = Coenonympha tullia california; Pelham, 2008, J. Res. Lepid. 40: 400

Coenonympha brenda; [LepIndex] ; [CoL]
Coenonympha california california; [NL4A], #1601a; [P&N23]; [iNaturalist] ; [BOA]
Coenonympha california; [NACL], #4586; [EBW]; [BOW]: pl. 18, f. 5; [NL4A], #1601; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 22; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
Coenonympha californica pulla; Dyar, 1903, Bull. U.S. nat. Mus. 52: 29
Coenonympha californica[sic, california]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 29
Coenonympha ochracea brenda; [NACL], #4584a; [NL4A], #1602a
Coenonympha tullia california; Pelham, 2008, J. Res. Lepid. 40: 400
californica[sic, california]; H. Edwards, 1881

Coenonympha amaryllis (Stoll, [1782])

S.Urals, Altai Mts, W.Siberia, E.Siberia, Transbaikalia, Amur, Ussuri, China, Mongolia, Korea. See [maps]

Papilio amaryllis Stoll, [1782]; in Cramer, Uitl. Kapellen 4 (32-32): 210, pl. 391, f. A, B; TL: "...de la Sibérie"
?Coenonympha amaryllis var. evanescens Alphéraky, 1889; in Romanoff, Mém. lép. 5: 118
?Coenonympha amaryllis var. ordossi Alphéraky, 1889; in Romanoff, Mém. lép. 5: 118
Coenonympha amaryllis v. accrescens Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 66
Coenonympha amaryllis borisovi Korshunov & Ivonin, 1996; (infrasubsp.) - Baikal region
- = Chortobius amaryllis accrescens; Korb, 2015, Eversmannia 43-44: 10

Chortobius amaryllis accrescens; Korb, 2015, Eversmannia 43-44: 9
Coenonympha amaryllis accrescens; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 f
Coenonympha amaryllis borisovi; [SZMN]
Coenonympha amaryllis emmonsi; [LepIndex]; [CoL] ; [iNaturalist]
Coenonympha amaryllis evanescens; Seitz, 1908, Gross-Schmett. Erde 1: 146
Coenonympha amaryllis forsteri; [LepIndex]; [CoL]
Coenonympha amaryllis ordossi; Seitz, 1908, Gross-Schmett. Erde 1: 146
Coenonympha amaryllis; [BRU], 194; Seitz, 1908, Gross-Schmett. Erde 1: 145; [BOW]: pl. 200, f. 5; [MRS], 403; [Otakar Kudrna]; Yakovlev, 2012, Nota lepid. 35 (1): 78; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [RFEL]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [SZMN]

400x398 (~41Kb) underside The valley of the Shivilig-Khem River just upstream its leaving the the southern foot of the East Tannu-Ola mounatin range for the Ubsu-Nur Hollow, Tes-Khem District, Tuva Republic, Siberia, Russia. 11th July 1990, Photo © Oleg Kosterin

600x800 (~94Kb) upperside A steppefied meadow on the Onon River right bank terrace at a pine forest edge, 5 km upstream of the village Nizhnii Tsasuchei, Onon District, Chita Province, Transbaikalia, Siberia, Russia, 30th June 1996 (Two individuals catched by a Thomisidae spider), Photo © Oleg Kosterin

C. a. amaryllis

Coenonympha amaryllis amaryllis; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 g; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [RFEL]; [iNaturalist]

C. a. rinda Ménétriés, 1859

amaryllis rinda Ménétriés, 1859; Bull. phys.-math. Acad. Sci. St. Pétersb. 17 (12-14): 217; TL: Amur

Coenonympha amaryllis rinda; Seitz, 1908, Gross-Schmett. Erde 1: 146, pl. 48 g; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [RFEL]

C. a. simingica Murayama

Coenonympha amaryllis simingica; [MRS], 403

Coenonympha rhodopensis Elwes, 1900

Italy, Romania, Bulgaria, Albania, N.Greece, Yugoslavia. See [maps] Eastern Large Heath

Coenonympha tiphon var. rhodopensis Elwes, 1900; Trans. ent. Soc. Lond. 1900 (2): 205; TL: Bulgaria, Rila Mts., Levi Iskar, 5000ft

Coenonympa iphis rhodopenses[sic, rhodopensis]; Seitz, 1908, Gross-Schmett. Erde 1: 147
Coenonympa tullia rhodopensis; [H&R], 214
Coenonympha rhodopensis; Abadjiev, 2000, Atalanta 31 (3/4): 461-466; [Otakar Kudrna]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]
rhodopensis; Abadjiev, 2002, Neue Ent. Nachr. 53: 27

The glycerion species-group

group Glycerion (Coenonympha); Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318

Coenonympha sunbecca Eversmann, 1843

Ghissar - Tarbagatai. See [maps]

Coenonympha sunbecca Eversmann, 1843; Bull. Soc. Imp. Nat. Moscou 16 (3): 538, pl. 7, f. 4a-b; TL: Zaisan, SE. Kazakhstan

Coenonympha sunbecca; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 498; Seitz, 1908, Gross-Schmett. Erde 1: 147, pl. 48 h i; [BRU], 197; [EBW]; [BOW]: pl. 200, f. 7; [MRS], 404; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

300x400 (~28Kb) underside A steppefied meadow in the middle flow of the Kora River, 30 km NE of the Tekeli City (1900 m above sea level), the North Dzhungarskiy Alatau mountain chain (NE part of Tien Shan), Taldy-Kurgan Province, SE Kazakhstan. 15th June, 1993 (on Artemisia santolinifolia Turcz. ex Besser), Photo © Oleg Kosterin

C. s. sunbecca

Coenonympha sunbecca sunbecca; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. s. alexandra Heyne, [1894]

sunbecca var. alexandra Heyne, [1894]; Palaearkt. Grossschm. 1: 624; TL: Kirghizsky Mts.
ptach Korb, 2005
- = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

Coenonympha sunbecca alexandra; Seitz, 1908, Gross-Schmett. Erde 1: 147; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

Coenonympha mahometana × sunbecca

?Coenonympha decolorata Wagner, 1913; Ent. Mitt. 2 (6): 189

Coenonympha mahometana × sunbecca; [BRU], 197
?Coenonympha decolorata; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47 (?)

Coenonympha mahometana Alphéraky, 1881

Tian-Shan. See [maps]

Coenonympha mahometana Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 428; TL: Kuldzha, Kunges Valley, W. China

Coenonympha iphis mahometana; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 d
Coenonympha mahometana; [BRU], 197; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [iNaturalist] ; [LEPIFORUM]

C. m. mahometana

Coenonympha mahometana mahometana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. m. acelae Hanus, 1996 Kirghizia

Coenonympha mahometana acelae Hanus, 1996; Atalanta 27 (1/2): 208; TL: Kirghiiza, Dolon Pass, 2800m

Coenonympha mahometana acelae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

Coenonympha glycerion (Borkhausen, 1788)

EU, S.Siberia, Japan. See [maps] Rostbraunes Wiesvögelchen Darrängefjäril Chestnut Heath Idänniittyperhonen Darrgräsfjäril Strzępotek glicerion

Papilio glycerion Borkhausen, 1788; Naturges. Eur. Schmett. 1: 90
Papilio iphis Denis & Shiffermüller, 1775; (preocc.)
Coenonympha iphis var. anaxagoras Assmus, 1857; Breslau Ent. Z. 10: 9
Papilio tiphon Esper, 1777; Die Schmett. Th. I, Bd. 1 (9): 341, (6): pl. 35, f. 3-4 (preocc.)
Coenonympha iphis var. carpathica Hormuzaki, 1897; Verh. zool.-bot. Ges. Wien. 47: 163
Coenonympha iphis anaxarete Fruhstorfer, 1910; Ent. Zs. 23 (53) 24 (1): 3; TL: Alpes Maritimes, Moulinet nr Menton
Coenonympha iphis ab. oikeia Dannehl, 1927; Mitt. Münch. Ent. Ges. 17 (1-6): 5

Coenonympha glycerion; [H&R], 221; [BOW]: pl. 5, f. 18; [MRS], 405; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BRU]; [Otakar Kudrna]; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SZMN] ; [SVF] ; [Mario Maier]
Coenonympha iphis; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 c; [SPRK]; [LepIndex] ; [CoL]
Coenonympha iphis ab. anaxagoras; Seitz, 1908, Gross-Schmett. Erde 1: 144

500x430 (~28Kb) underside Finland, Ylöjärvi, Teivaala, 6828:320, 16.7.1996, Photo © Tero Piirainen	800x540 (~51Kb) underside Finland, Vihti, Nummela, ?July 2002, Photo 10072002_24 © Timo Viitanen	470x443 (~80Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý
470x379 (~76Kb) underside Czech Republic, Blansko, 22.7.2004, Photo © Michal Koupý	600x400 (~60Kb) underside Estonia, Hiiumaa, Kootsaare nina. 24.07.2004, Photo © Hannu Tanner	724x724 (~116Kb) underside Russia, Moscow area, 10.06.2007, Photo © D. Smirnov
710x522 (~96Kb) ♂ underside Russia: Moscow area (36°25'E, 56°00'N), 14.7.2008, Photo © D. Smirnov	811x660 (~113Kb) ♀ underside Russia, Moscow area, 18.8.2010 (36°25'E, 56°00'N), Photo © D. Smirnov	333x278 (~15Kb) ♀
320x275 (~23Kb) ♂

C. g. glycerion

Coenonympha glycerion glycerion; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [iNaturalist] ; [SZMN]

C. g. iphicles (Staudinger, 1892) N.Alai, Sayan, Transbaikalia, Amur, Ussuri

Coenonympha iphis var. iphicles Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 338; TL: Kentei
Coenonympha iphis var. heroides Christoph, 1893; Dt. Ent. Z. Iris 6 (1): 87

Coenonympha glycerion heroides; [RFEL]; [SZMN]
Coenonympha glycerion iphicles; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; Yakovlev, 2012, Nota lepid. 35 (1): 78; [iNaturalist] ; [SZMN]
Coenonympha iphis iphicles; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 c

872x1120 (~262Kb) underside A long-fallow land on the Akkem River right bank terrace at the Oroktoi Brook mouth, Katunskii Range southern foot, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Republic, West Siberia, Russia. 8th July 2007, Photo © Oleg Kosterin

979x1081 (~180Kb) underside Russia, West Siberia, the Altai Mts., Altai Republic, Shebalino District, the Ulus-Cherga village environs, a herbaceous meadow on southern slope. 11th July 2007, Photo © Oleg Kosterin

C. g. iphina Staudinger, 1892 Saur, Tarbagatai

Coenonympha iphis var. iphina Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 339; TL: Tarbagatai

Coenonympha glycerion iphina; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47
Coenonympha iphis ab. iphina; Seitz, 1908, Gross-Schmett. Erde 1: 144

C. g. wutaica Murayama

Coenonympha glycerion wutaica; [MRS], 405

C. g. alta Sheljuzhko, 1937 Caucasus

glycerion alta Sheljuzhko, 1937; Festschr. Embrik Strand, Riga 2: 352

Coenonympha glycerion alta; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47
Coenonympha iphis alta; [LepIndex]; [CoL]

C. g. korshunovi Nekrutenko, 1978 Crimea

glycerion korshunovi Nekrutenko, 1978; 646

Coenonympha amyntas korshunovi; [LepIndex]; [CoL]
Coenonympha glycerion korshunovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. g. beljaevi Dubatolov, 1997 S.Primorye

Coenonympha glycerion beljaevi Dubatolov, 1997; Far Eastern Ent. 44: 8; TL: S.Primorye, Spassk Districk

Coenonympha glycerion beljaevi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [RFEL]; [SZMN]

Coenonympha iphioides Staudinger, 1870

Spain. See [maps] Spanish Heath Castaneo morena

Coenonympha iphis var. iphioides Staudinger, 1870; Berl. Ent. Z. 14: 101
Coenonimpha[sic] iphioides pseudoamyntas de Sagarra, 1930; Butll. Inst. catal. Hist. nat. (2) 10 (7): 113

Coenonympha glycerion iphioides; [iNaturalist]
Coenonympha iphioides; [H&R], 222; [BOW]: pl. 6, f. 2; [Otakar Kudrna]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LEPIFORUM] ; [MBE] ; [MBE]
Coenonympha leander iphioides; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 c; [LepIndex]; [CoL]

The hero species-group

group Hero (Coenonympha); Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318

Coenonympha corinna (Hübner, 1804)

Corsica, Sardinia, Elba. See [maps] Corsican Heath

Papilio corinna Hübner, 1804; Samml. eur. Schmett. [1]: 40, pl. 105, f. 536, 537
?Coenonympha corinna corinnaeformis Verity, 1914; Boll. Soc. ent. ital. 45: 228

Coenonympha corinna trettaui; [LepIndex]; [CoL] ; [iNaturalist]
Coenonympha corinna; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 f; [H&R], 216; [BOW]: pl. 5, f. 16; [Otakar Kudrna]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]

Coenonympha elbana Staudiger, 1901 Elba, Giglio, Monte Argentario

Coenonympha elbana Staudiger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 66

Coenonympha corinna elbana; Seitz, 1908, Gross-Schmett. Erde 1: 145
Coenonympha elbana; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LEPIFORUM] ; [MBE] ; [MBE]

Coenonympha arcanioides (Pierret, 1837)

Morocco, Algeria, Tunisia. See [maps] Moroccan Pearly Heath

Satyrus arcanioides Pierret, 1837; Ann. Soc. ent. Fr. 6: 306; TL: Province of Oran (Algeria)
?Satyrus marginalis Butler, 1868; Cat. diurn. Lep. Satyridae Brit. Mus.: 42
- = ?Coenonympha arcania;
Coenonympha arcanioides f. major Seitz, 1908; Gross-Schmett. Erde 1: 143, pl. 48[e]; TL: Algeria, Morocco
Coenonympha arcanioides gen. aest. holli Oberthür, 1910; Ètud. Lépid. Comp. 4: 20; TL: Algeria
Coenonympha arcanioides ab. biocellata Oberthür, 1910; Ètud. Lépid. Comp. 4: 21; TL: [Algeria]
Coenonympha arcanioides f. pluriocellata Stetter-Stättermayer, 1933; Int. Ent. Zs. 27 (30): 340; TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. inocellata Stetter-Stättermayer, 1933; Int. Ent. Zs. 27 (30): 340; TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. obliterata Stetter-Stättermayer, 1933; Int. Ent. Zs. 27 (30): 340; TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. nigro-ocellata Stetter-Stättermayer, 1933; Int. Ent. Zs. 27 (30): 340; TL: Bône [Annaba] (Algeria)

Coenonympha arcanioides; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 e; [H&R], 220; [BOW]: pl. 5, f. 12; [BMAT]: 88, pl. 28, f. 45-49; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]

Coenonympha vaucheri Blachier, 1905

Morocco. See [maps] Vaucher's Heath

Coenonympha vaucheri Blachier, 1905; Bull. Soc. ent. Fr. 1905: 213; TL: High Atlas Mts. (Morocco)

Coenonympha vaucheri; Seitz, 1908, Gross-Schmett. Erde 1: 145, pl. 48 e; [H&R], 218; [BOW]: pl. 115, f. 6; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [MBE] ; [MBE]

C. v. vaucheri Morocco (High Atlas)

Coenonympha vaucheri ab. geminipuncta Blachier, 1905; Bull. Soc. ent. Fr. 1905: 213; TL: High Atlas Mts. (Morocco)
Coenonympha meade-waldoi Elwes, 1905; Trans. ent. Soc. Lond. 1905 (4): 377, pl. 19, f. 1-2
Coenonympha vaucheri f. mediocellis Le Cerf, 1923; Bull. Soc. ent. Fr. 28 (17): 224; TL: Grand Atlas, Djebel Tachdirt, 3100-3250m

Coenonympha meadewaldoi; [LepIndex] ; [CoL]
Coenonympha vaucheri vaucheri; [BMAT]: 86, pl. 29, f. 32-40

C. v. annoceuri Wyatt, 1952 Morocco (Middle Atlas)

Coenonympha vaucheri annoceuri Wyatt, 1952; Zs. Wiener ent. Ges. 37: 175, pl. 21, f. B1-B4; TL: Annoceur (Morocco)

Coenonympha vaucheri annoceuri; [BMAT]: 87, pl. 29, f. 41-48; [LepIndex]; [CoL]

C. v. rifensis Weiss, 1979 Morocco (W.Rif Mts.)

Coenonympha vaucheri rifensis Weiss, 1979; Entomops 48: 271, f. 1; TL: Dj. Lakraa (Morocco)

Coenonympha vaucheri rifensis; [BMAT]: 87, pl. 29, f. 49-57; [LepIndex]; [CoL]

C. v. beraberensis Lay & Rose, 1979 Morocco (Tizi-n'Ouguerd-Zegzaoune, Dj.Aourach, Imlil, Lake Tislit, etc.)

Coenonympha vaucheri beraberensis Lay & Rose, 1979; Ent. Zs. 89 (13): 143, f. 2; TL: [Tizi-n'Ouguerd-Zegzaoune] (Morocco)

Coenonympha vaucheri beraberensis; [BMAT]: 87, pl. 29, f. 58-66; [LepIndex]; [CoL]

Coenonympha arcania (Linnaeus, 1761)

EU, Asia Minor, S.Russia, S.Urals. See [maps] Mancha Leonada Pearly Heath Pärlgräsfjäril Strzępotek perełkowiec Perlgrasfalter

Papilio arcania Linnaeus, 1761; Fauna Suecica (Edn 2): 273; TL: Sweden
Papilio amyntas Poda, 1761; Ins. Mus. Graecensis: 79
?insubrica Frey, 1882; Mitt. schweiz. ent. Ges. 6 (7): 353
?melania Oberthür
?badensis Reutti
Coenonympha arcania balestrei Fruhstorfer, 1910; Ent. Zs. 23 (53) 24 (1): 3
Coenonympha arcania saleviana Fruhstorfer, 1910; Ent. Zs. 23 (53) 24 (1): 3; TL: Savoy, Switzerland
Coenonympha arcania chrysoaspida Fruhstorfer, 1910; Ent. Zs. 23 (53) 24 (1): 4; TL: ;Saratow
Coenonympha arcania triumphans Fruhstorfer, 1910; Soc. ent. 25 (14): 55
Coenonympha arcanius tenuelimbo Verity, 1914; Boll. Soc. ent. ital. 45: 223, pl. 1, f. 31-33
Coenonympha arcania clorinda de Sagarra, 1924; Butll. Inst. Catal. Hist. nat. (2) 4 (9): 199; TL: "Orihuela del Tremedal"
Coenonympha arcania ab. euthymia Dannehl, 1927; Mitt. Münch. Ent. Ges. 17 (1-6): 4

Coenonympha amyntas; [LepIndex] ; [CoL]
Coenonympha arcania baqueroensis; [LepIndex]; [CoL]
Coenonympha arcania momos; [LepIndex]; [CoL]
Coenonympha arcania salomonis; [LepIndex]; [CoL]
Coenonympha arcania satyrionides; [LepIndex]; [CoL]
Coenonympha arcania skypetarum; [LepIndex]; [CoL]
Coenonympha arcania; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 d; [H&R], 219; [BRU], 195; [EBW]; [BOW]: pl. 5, f. 15; [MRS], 405; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF] ; [Mario Maier]
Coenonympha arcania ab. badensis; Seitz, 1908, Gross-Schmett. Erde 1: 144
Coenonympha arcania ab. insubrica; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 d
Coenonympha arcania ab. melania; Seitz, 1908, Gross-Schmett. Erde 1: 144

500x471 (~45Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes	652x452 (~27Kb) ♂ underside Poland, Tresta near Tomaszow Mazowiecki, 07.1997, Photo © Marek Michalski leg.	309x300 (~24Kb) underside An opening in an oak forest, the town Krymsk environs, Krymsk District, Krasnodarskii Krai Province, S Russia. 26th May 1990, Photo © Oleg Kosterin
517x689 (~57Kb) underside Sweden, Ekebo, Julita, 5.7.2003, Photo © Leif Wahlberg	479x639 (~66Kb) underside Sweden, Ekebo, Julita, 20.6.2004, Photo © Leif Wahlberg	470x441 (~93Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý
590x620 (~88Kb) underside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov

C. a. arcania

Coenonympha arcania arcania; [H&R], 219; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

C. a. caucasica Jachontov, 1914

arcania caucasica Jachontov, 1914; Revue Russe Ent. 14: 298

Coenonympha arcania caucasica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

Coenonympha gardetta (de Prunner, 1798)

Alps, N.Balkans. See [maps] Alpine Heath

Papilio gardetta de Prunner, 1798; Lepid. Pedemont.: 74
- = Coenonympha pamphilus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 30
Papilio satyrion Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 24, pl. 122, f. 2
- = ; [H&R], 220
Papilio philea Hübner, [1800]; Samml. eur. Schmett. [1]: pl. 53, f. 254-255
- = ; [H&R], 220
?Papilio neoclides Hübner, 1796; Samml. eur. Schmett. [1]: 41, f. 254-255 (philea)
neoclides (Hübner, 1805)
- = ; [H&R], 220

Coenonympha arcania satyrion; Seitz, 1908, Gross-Schmett. Erde 1: 144, pl. 48 d
Coenonympha gardetta gardetta; [iNaturalist]
Coenonympha gardetta; [H&R], 220; [Otakar Kudrna]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [Mario Maier]
Coenonympha neoclidis[sic, neoclides?]; Hübner, [1819], Verz. bek. Schmett. (5): 65

Coenonympha semenovi Alphéraky, 1887

-. See [maps]

Coenonympha semenovi Alphéraky, 1887; in Romanoff, Mem. lép. 3: 405
Coenonympha semenovi vera Kocman, 1996; Lambillionea 96: 41; TL: China, N.Qinghai, East Chilien Shan, Papao
- = ; Lang, 2018, Atlanta 4 (1-4): 115

Coenonympha semenovi; Seitz, 1908, Gross-Schmett. Erde 1: 147, pl. 48 i; [MRS], 404; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); Lang, 2018, Atlanta 4 (1-4): 115; [LepIndex] ; [CoL] ; [iNaturalist]
?Coenonympha semenori[sic, semenovi]; [BOW]: pl. 200, f. 6

C. s. semenovi Qinghai, NE.Tibet

Coenonympha semenovi semenovi; Lang, 2018, Atlanta 4 (1-4): 115

C. s. leanotchka Hemming, 1933 SW.Gansu, E.Qinghai, Sichuan

Coenonympha semenovi leanotchka Hemming, 1933; Entomologist 66 (12): 275
Coenonympha semenovi var. obscura Alphéraky, 1897; in Romanoff, Mém. Lép. 9: 111; TL: Ta-tsien-lu (preocc. _Coenonympha leander var. obscura_ Rühl, 1894)
- = ; Lang, 2018, Atlanta 4 (1-4): 115
Coenonympha semenovi szechwana Bang-Haas, 1934; Ent. Zs. 48 (14): 110; TL: W.China, Tatsienlu, Putsufong, Howkow, 7000-10000
- = ; Lang, 2018, Atlanta 4 (1-4): 115
Coenonympha semenovi arnoldi Bang-Haas, 1934; Ent. Zs. 48 (14): 110; TL: Kuku nor mer. or. Alt Tau, Hsi king shan, 2500m
- = ; Lang, 2018, Atlanta 4 (1-4): 115

Coenonympha semenovi arnoldi; [LepIndex]; [CoL]
Coenonympha semenovi leanotchka; Lang, 2018, Atlanta 4 (1-4): 115
Coenonympha semenovi szechwana; [LepIndex]; [CoL]

C. s. jiadengyuica Huang & Murayama, 1992 Xianjing

Coenonympha semenovi jiadengyuica Huang & Murayama, 1992; Tyô to Ga 43 (1): 5, f. 15; TL: Jiadengyu, Altai, 1380m

Coenonympha semenovi jiadengyuica; [MRS], 404

C. s. sala Kocman, 1995 Qinghai

Coenonympha semenovi sala Kocman, 1995; Lambillionea 95: 67; TL: Chaka-Wulan, Qinghai

Coenonympha semenovi sala; Lang, 2018, Atlanta 4 (1-4): 115

C. s. goateri Lang, 2018 Sichuan

Coenonympha semenovi goateri Lang, 2018; Atlanta 4 (1-4): 116; TL: Sichuan, Lixian, Miyalo, 3700-3800m

C. s. yufei Lang, 2018 Shaanxi

Coenonympha semenovi yufei Lang, 2018; Atlanta 4 (1-4): 116; TL: Shaanxi, Huxian, Zhuque, Bingjing-ding, 2800-300m

Coenonympha hero (Linnaeus, 1761)

CEU, NEU, Temperate Asia, Amur, Korea, Japan. See [maps] Madängefjäril Strzępotek hero Brun gräsfjäril Scarce Heath Ruskoniittyperhonen Wald-Wiesenvögelchen

Papilio hero Linnaeus, 1761; Fauna Suecica (Edn 2): 274; TL: Sweden
?stolida Schilde
?Coenonympha hero var. areteoides Fologne, 1874; Ann. Soc. ent. Belg. 17 (C. R.): xcix

Coenonympha hero angustefasciata; [LepIndex]; [CoL]
Coenonympha hero; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 b; [H&R], 223; [BRU], 196; [EBW]; [BOW]: pl. 6, f. 1; [MRS], 404; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [RFEL]; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF] ; [Mario Maier]
Coenonympha hero ab. stolida; Seitz, 1908, Gross-Schmett. Erde 1: 143
?Coenonympha hero ab. areteoides; Seitz, 1908, Gross-Schmett. Erde 1: 143

C. h. hero

Coenonympha hero hero; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47

?C. h. sabaeus Fabricius, 1775 E.Europe, Urals, W.Siberia

C. h. perseis Lederer, 1853 up to Pacific

Coenonympha hero var. sibirica Fuchs, 1899; Jb. Nassau Ver. Nat. 52: 126

Coenonympha hero perseis; [MRS], 404; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 47; [RFEL]; [iNaturalist]
Coenonympha hero ab. perseis; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 a

478x640 (~92Kb) ♂ upperside underside Russia, N Tuva, Piykhemskiy distr., h=800., 06.1997 Vashchenko S.N. leg, Photo © D. Smirnov

C. h. neoperseis Fruhstorfer, 1908 Japan

Coenonympha hero neoperseis Fruhstorfer, 1908; Int. ent. Zs. 2 (2): 10; TL: Japan, nr. Sapporo

Coenonympha hero neoperseis; Kudrna, 1974, Atalanta 5: 100; [iNaturalist]
Coenonympha hero ab. neoperseis; Seitz, 1908, Gross-Schmett. Erde 1: 143

C. h. latifasciata Matsumura, 1925 Hokkaido

Coenonympha heros[sic] latifasciata Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 94; TL: Ohtsu, Hokkaido

Coenonympha hero latifasciata; Kudrna, 1974, Atalanta 5: 100; [iNaturalist]

C. h. pilwonis Matsumura, 1925 Sakhalin

Coenonympha heros[sic] pilwonis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 95, pl. 8, f. 2 ♂; TL: Saghalin

C. h. coreana Matsumura, 1927 Korea

Coenonympha hero coreana Matsumura, 1927; Insecta Matsumurana 1 (4): 164; TL: Corea

Coenonympha leander (Esper, 1784)

Hungary, Bulgaria, S.Russia, Asia Minor, Armenia, Iran. See [maps] Russian Heath

Papilio leander Esper, 1784; Die Schmett. Th. I, Bd. 2 (9): 176, pl. 89, f. 5; TL: Volga region, Russia

Coenonympha leander gallica; [LepIndex]; [CoL]
Coenonympha leander katarae; [LepIndex]; [CoL]
Coenonympha leander; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 c; [H&R], 221; [BRU], 195; [BOW]: pl. 6, f. 3; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; Kodandaramaiah & Wahlberg, 2009, Syst. Ent. 34: 318 (note); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]

C. l. leander

Coenonympha leander leander; [H&R], 221; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; [iNaturalist]

C. l. obscura Rühl, 1894 Caucasus Minor, Armenian Highland, Talysh

leander obscura Rühl, 1894; Palaearkt. Grossschm. 1: 607

Coenonympha leander obscura; Seitz, 1908, Gross-Schmett. Erde 1: 143, pl. 48 c; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 48; [LepIndex]; [CoL] ; [iNaturalist]

C. l. iranica Schwingenschuss, 1939

Coenonympha leander iranica Schwingenschuss, 1939; Ent. Zs. 53 (2): 13, (8):62; TL: Kendevan

Coenonympha leander iranica; [LepIndex]; [CoL]

C. l. wageneri Gross & Ebert, 1975

Coenonympha leander wageneri Gross & Ebert, 1975; J. ent. Soc. Iran, suppl. 1: 26; TL: E.Azerbaijan/Glian: Paß der Staße Firuz-Abad-Hashtpar, 2300m

C. l. dierli Koçak, 1977 Turkey

Coenonympha leander dierli Koçak, 1977; Atalanta 8 (2): 131; TL: Demirkazik

Coenonympha leander dierli; [LepIndex]; [CoL]

Coenonympha leander trevincae Wyatt, 1952; Zs. Wiender ent. Ges. 37: 207; TL: Peña Trevinca, Orense, NW.Spain, 1560m

Coenonympha pamphilus ab. caeca Oberthür, 1910; Ètud. Lépid. Comp. 4: 48
Coenonympha pamphilus ab. caecaella Strand, 1927; Arch. Naturg. 91 A (12): 281 (repl. Coenonympha pamphilus ab. caeca Oberthür, 1910)

Coenonympha caeca; [CoL]

Coenonympha tiphon italica Verity, 1914; Boll. Soc. ent. ital. 45: 222, pl. 1, f. 24-27

Coenonympha tiphon var. grisescens Christoph, 1893; Dt. Ent. Z. Iris 6 (1): 87

References:

[A. Timchenko]
Andrew Timchenko; Beijing, China (ex-Moscow, Russia)
[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BHL] The
Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BnF]
Gallica / Bibliothèque nationale de France; Annales de la Société entomologique de France; Bulletin de la Sociéte entomologique de France
[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[ESP] :ESP#c Heppner, 1981
[EVERSMANNIA]
Eversmannia contents
[Enrique Garcia-Barros]
Enrique García-Barros; Dept. Biol. (Zool.); Univ. Autonoma de Madrid
[FEE]
Far Eastern Entomologist
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Hannu Tanner]
Hannu Tanner, Siuntio, Finland; e-mail: Hannu Tanner; http://www.kolumbus.fi/hannu.j.tanner/coleoptera/
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[Jean Delacre]
http://homepage.mac.com/jdelacre
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[Leif Wahlberg]
Julita, Sweden; e-mail: Leif Wahlberg
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MPG]
North American Moth Photographers Group
[Marek Michalski] Lodz, Poland
e-mail: Marek Michalski
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Maurizio Elia]
Ravenna, Italy; e-mail: Maurizio Elia
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michal Koupý]
Czech Republic
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler] Opler & Warren, 2003
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[P&N23] Pohl & Nanz, 2023
Annotated Taxonomic Checklist of the Lepidoptera of North America, north of Mexico
[PERSEE]
Annales de la Société linnéenne de Lyon; Bulletin mensuel de la Société linnéenne de Lyon; Publications de la Société Linnéenne de Lyon; Bulletin de la Société entomologique de France
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN] Lepidoptera
Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Szabolcs Safian]
Hungary; Hungarian Lepidopterologial Society; Butterflies of Ghana
[Tero Piirainen] Tampere, Finland
e-mail: Tero Piirainen
[Timo Viitanen]
http://www.kuvarkisto.com/
[USGS] Northern Prairie Wildlife Research Center
http://www.npwrc.usgs.gov/index.htm
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Abadjiev, 2000
Type of Balkan butterflies. II. Eastern Large Heath, Coenonympha rhodopensis Elwes, 1900: type locality and lectotype designation Atalanta 31 (3/4) : 461-466
Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 53 : 3-53
Alphéraky, 1881
Lépidoptères du district de Kouldjà et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)
Alphéraky, 1887
Diagnoses de quelques lépidoptères inédits du Thibet in Romanoff, Mem. lép. 3 : 403-406
Alphéraky, 1888
Neue Lepidopteren Stettin Ent. Ztg 49 : 66-69
Alphéraky, 1889
Lépidoptères rapportés de ia Chine et de la Mongolie par G. N. Potanine in Romanoff, Mém. lép. 5 : 90-123, pl. 5
Alphéraky, 1897
Lépidoptéres des provinces chinoises Sé-Tchouen et Kam recueillis, en 1893, par M-r G. N. Potaine in Romanoff, Mém. Lép. 9 : 83-149
Alphéraky, 1897
Lepidopteren aus Kamtschatka gesammelt von O. Herz, bearbeitet von S. Alpheraky in Romanoff, Mém. Lép. 9 : 301-347
Assmus, 1857
Coenonympha anaxagoras eine neue Varietät der Coenonympha iphis und ein Beitrag zur Naturgeschichte von Larentia Pyropata Hüb. Breslau Ent. Z. 10 : 9-10
Bang-Haas, 1927
Horae Macrolepidopterologicae Regionis Palaearcticae Horae Macrolep. Palaearct. 1 : 1-128, pl. 1-11
Bang-Haas, 1934
Neubeschreibungen und Berichtigungen der Palaearktischen Macrolepidopterenfauna. VIII-XVII Ent. Zs. 48 (1): 7-8, (2): 16, (6): 48, (7): 56, (9): 71-72, (14): 108-110, (16): 128, (17): 135, (19): 152 (1935), (23): 183-184 (1935)
Barnes & Benjamin, 1926
Notes on diurnal Lepidoptera, with addition and corrections to the recent "List of Diurnal Lepidoptera" Bull. South. Calif. Acad. Sci. 25 (3) : 88-98
Blachier, 1905
Descriptions de Lépidoptères nouveaux de Maroc Bull. Soc. ent. Fr. 1905 : 212-215
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Borkhausen, 1788
Naturgeschichte der europäischen Schmetterlinge nach Systematischer Ordnung. 1 Naturges. Eur. Schmett. 1 : 3-288, pl. [1]
Butler, 1866
Descriptions of some new species of diurnal Lepidoptera in the collection of the British Museum Ent. mon. Mag. 3 (28) : 76-78
Butler, 1868
A catalogue of diurnal Lepidoptera of the family Satyridae in the collection of the British Museum Cat. diurn. Lep. Satyridae Brit. Mus. : 1-211, pl. 1-5
Butler, 1877
On Rhopalocera from Japan and Shanghai, with Descriptions of new Species Ann. Mag. nat. Hist. (4) 19 (109) : 91-97
Le Cerf, 1923
Lépidoptères nouveaux du Maroc Bull. Soc. ent. Fr. 28 (17) : 224
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
Christoph, 1893
Lepidoptera Nova Faunae Palaearcticae Dt. Ent. Z. Iris 6 (1) : 86-96
Comstock, 1925
Studies in Pacific Coast Lepidoptera Bull. South. Calif. Acad. Sci. 24 (1): 3-4, (2): 35-37, pl. 2, (3): 62-68
Dannehl, 1927
Neue Formen und geographische Rassen aus meinen Rhopalocecer-Ausbeuten der letzten Jahre Mitt. Münch. Ent. Ges. 17 (1-6) : 1-8
Davenport, 1936
A new American Coenonympha (Lepid. Satyridae) Can. Ent. 68 (4) : 79-80
Davenport, 1941
The Butterflies of the Satyrid Genus Coenonympha Bull. Mus. comp. Zool. Harv. 87 (4) : 215-349, pl. 1-10
dos Passos, 1935
Some butterflies of southern Newfoundland with descriptions of new subspecies Can. Ent. 67 (4) : 82-88
Doubleday, 1859
Zoologist Synonymic List of British Butterfles and Moths Zool. Syn. List Br. Butts Moths (edn 2) : 1-40
Westwood, [1851]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 18*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29 (15-23) (1848), (1): pl. 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Dubatolov, 1997
New data on taxonomy of Lycaenidae, Nymphalidae and Satyridae (Lepidoptera, Rhopalocera) of the Asian part of Russia Far Eastern Ent. 44 : 1-12
Dunning & Pickard, 1858
An Accentuated List of the British Lepidoptera Accent. List Brit. Lep.: : 1-118
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1861
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1869
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 (4) : 369-376
Edwards, 1871
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (3/4) : 205-216
Edwards, 1872
List of species of butterflies collected by Campbell Carrington and Wm. B. Logan, of the expedition [to Montana], in 1871. Preliminary report [5th Ann. rep.] U. S. Geological Survey of Montana, by F. V. Hayden in Hayden, Geol. Surv. Montana: 466-467
Edwards, 1881
Coenonympha elko Can. Ent. 13 (3) : 57-58
H. Edwards, 1877
Pacific Coast Lepidoptera. - No. 22. Notes on some Diurnal Lepidoptera, with description of new varieties Proc. Calif. Acad. Sci. 7 (12) : 163-173
H. Edwards, 1881
On some apparently new forms of diurnal lepidoptera Papilio 1 (4) : 50-55
Elwes, 1900
On the Butterflies of Bulgaria Trans. ent. Soc. Lond. 1900 (2) : 183-206, pl. 4
Erschoff, 1874
Travels in Turkestan. Volume 2. Zoogeographical Investigations. Lepidoptera [in Russian] in Fedschenko, Travels in Turkestan. 2 (5) : 1-128, pl. 1-6
Esper, 1777
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)
Esper, 1780; Esper, 1782; Esper, 1784
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Esper, 1794
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])
Esper, 1805
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 2 Die Schmett., Th I, Suppl. Th. 2 (11): 1-24, pl. 117-122 (1805), (12): 25-48, pl. 123-126 [1830]
Eversmann, 1843
Quaedam lepidopterorum species novae in montibus Uralensibus et Altaicus habitantes nunc descriptae et depictae Bull. Soc. Imp. Nat. Moscou 16 (3) : 535-553, pl. 7-10
Eversmann, 1851
Description de quelques nouvelles espéce de Lépidoptéres de la Russie Bull. Soc. imp. Nat. Moscou 24 (2) : 610-644
Fabricius, 1777
Genera Insectorum eorumque characters naturales secundum numerum, figuram, situm et proportionem ... Genera Insectorum : 1-310
Fabricius, 1787
Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus Mantissa Insectorum 2 : 1-382
Field, 1937
A new seasonal form of Coenonympha ampelos Edwards (Lepid.: Satyridae) Can. Ent. 69 (11) : 249-250
Frey, 1882
Zweiter Nachtrag zur Lepidopteren-Fauna der Schweiz Mitt. schweiz. ent. Ges. 6 (7) : 349-375
Freyer, 1845
Neuere Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur. (65-80) Neuere Beitr. Schmett. 5 : 1-40 (1842), : 41-92 (1844), : 93-164 (1845)
Fruhstorfer, 1908
Neue palaearktische Satyriden Int. ent. Zs. 2 (2) : 9-10 (11 April)
Fruhstorfer, 1910
Neue palaearctische Satyriden Ent. Zs. 23 (53) 23 (53): 239-240 (2 April), 24 (1): 3-4 (9 April)
Fruhstorfer, 1910
Neue palaearktische Rhopaloceren Soc. ent. 25 (13): 50-52 (24 September, 1910), (14): 54-55 (8 October, 1910), (15): 58-60, f. 1-4 (22 October, 1910), (24): 95-96 (25 February, 1911)
Fuchs, 1899
Macrolepidopteren der Loreley-gegend un verwandte formen Jb. Nassau Ver. Nat. 52 : 117-158
Gross & Ebert, 1975
Beiträge zur Kenntnis der Rhopaloceren Irans - Neue Taxa der Pieridae, Satyridae und Nymphalidae J. ent. Soc. Iran, suppl. 1 : 8-45
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Gussich, 1917
Ein Beitrag zur Schmetterlingsfauna Kroatiens Godina 29 : 209-225
Hanus, 1996
New butterfly taxa from Kirghizia (Lepidoptera, Rhopalocera) Atalanta 27 (1/2) : 203-210
Hemming, 1933
Holarctic butterflies: miscellaneous notes on nomenclature Entomologist 66 (12) : 275-279
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Holland, 1900
Alaska Insects Ent. News 11 (3): 381-389 (part I), (4): 416-423 (part II)
Hormuzaki, 1897
Die Schmetterlinge (Lepidoptera) der Bukowina Verh. zool.-bot. Ges. Wien. 47 : 70-103, : 120-168, : 233-246, : 312-341
Huang & Murayama, 1992
Butterflies of Xinjiang province, China Tyô to Ga 43 (1) : 1-22
Hübner, [1790-1793]
Der Schmetterlinge Lepidoptera Linnei, Europäisches Heer, [1790-1793] Schmett. Lep. linn.: [1-3], 4-16
Hübner, 1796; Hübner, [1800]; Hübner, 1804
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 982-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Jachontov, 1914
La faune lépidoptèrologique russe et les types de l'Europe centrale [in Russian] Revue Russe Ent. 14 : 295-306
Koçak, 1977
New Lepidoptera from Turkey V Atalanta 8 (2) : 126-147
Kocman, 1995
Some new subspecies of the genus Parnassius, Boloria and Coenonympha from China. (Lepidoptera: Papilionidae, Nymphalidae and Satyridae) Lambillionea 95 : 63-67
Kocman, 1996
Some new subspecies of the genus Parnassius and Coenonympha from China (Lepidoptera: Papilionidae, Satyridae) Lambillionea 96 : 37-42
Kodandaramaiah & Wahlberg, 2009
Phylogeny and biogeography of Coenonympha butterflies (Nymphalidae: Satyrinae) - patterns of colonization in the Holarctic Syst. Ent. 34 : 315-323
Kollar, [1849]
Ueber den Charakter der Insecten-Fauna von SüdPersien Denkschr. Akad. Wiss. Wien. 1 : 42-53
Korb, 2015
To the systematics and nomenclature of palaearctic Satyridae (Lepidoptera) Eversmannia 43-44 : 8-10
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Krulikowsky, 1908
Einige neue Varietäten und Aberrationen der Lepidopteren des östlichen Russlands Soc. Ent. 23 (1): 2-3, (2): 11-12, (3): 18
Krzywicki, 1967
Papilionoidea und Hesperioidea (Lepidoptera) des Bialowiezaer Urwalds Ann. zool. Warsz. 25 (1) : 1-213
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Lang, 2018
Notes on Coenonympha semenovi Alphéraky, 1887 with description of two new subspecies (Lepidoptera, Nymphalidae & Satyrinae) Atlanta 4 (1-4) : 115-120
Lederer, 1870
Contributions a la faune des Lepidopteres de la Transcaucasie Ann. Soc. ent. Belg. 13 : 17-54, pl. 1-2
Leech, [1892]
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-296, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
[BOW]; Lewis, 1974
Butterflies of the World
Lingonblad, 1950
Neue Schmetterlingsformen Notulae Ent. 29 (3-4) : 79-82
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1761
Fauna Suecica Sistens Animalia Sueciae Regni: Mamalia, Aves, Amphibia, Pisces, Insecta, Vermes. Distributa per Classes, Ordines, Genera, Species, cum differentiis Specierum, Synonymis Auctorum, Nominibus Incolarrum, Locis Natalium, Descriptionibus Insectorum Fauna Suecica (Edn 2) : 1-578
Matsumura, 1925
An enumeration of the butterflies and moths from Saghalien, with descriptions of new species and subspecies J. Coll. Agric. Hokkaido imp. Univ. 15 : 83-196, pl. 8-11
Matsumura, 1927
A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170
McDunnough, 1928
Notes on Canadian diurnal Lepidoptera Can. ent. 60 (11) : 266-275
McDunnough, 1939
A new Coenonympha race from northeastern New Brunswick Can. ent. 71 (12): 266
Elwes in Meade-Waldo, 1905
On a collection of Butterflies made in Morocco, in 1900-01-02 Trans. ent. Soc. Lond. 1905 (4) : 369-381, pl. 18-19
Ménétriés, 1859
Lépidoptères de la Sibérie orientale et en particulier des rives de l'Amour Bull. phys.-math. Acad. Sci. St. Pétersb. 17 (12-14) : 212-221, Mélanges biol. St.-Pétersb. 3 (1): 99-113 (1858)
Müller, 1764
Fauna Insectorum Fridrichsdalina sive Methodica Descriptio Insectorum Agri Fridrichsdalensis cum Characteribus Genericis et Specificis, Nominibus Trivialibus, Locis Natalibus, Iconibus Allegatis, Novsque Pluribus Speciebus Additis Fauna Ins. Fridr. : 1-96
Munguira, García-Barros & Martín, 1997
Plantas nutricias de los licénidos y satirinos españoles (Lepidoptera: Lycaenidae y Nymphalidae) Boletín de la Asociación española de Entomología, 21 : 29-53
Oberthür, 1874
Lépidoptères nouveaux d'Algérie Petites Nouv. Ent. 1 (103) : 412-413
Oberthür, 1881
Lépidoptères d'Algerie Étud. d'Ent. 6 : 45-96
Oberthür, 1910
Notes pour servir à établir la Faune Française et Algérienne des Lépidoptères Ètud. Lépid. Comp. 4 : 15-691, pl. 35-58
Oberthür, 1922
Les Lépidoptères du Maroc Étud. Lépid. Comp. 19 (1): 13-402, (2): pl. 530-546
Pelham, 2008
A catalogue of the butterflies of the United States and Canada J. Res. Lepid. 40 : 1-652
Pierret, 1837
Description de deux nouvelles espèces du genre Satyre (Latr.) Ann. Soc. ent. Fr. 6 : 303-307, pl. 12
Poda, 1761
Insecta Musei Graecensis, quae in ordines, genera et species juxta Systema Naturae Caroli Linnaei digessit Ins. Mus. Graecensis: 8+127, 2pls
Porter & Mattoon, 1989
A new subspecies of Coenonympha tullia (Müller) (Nymphalidae: Satyrinae) confined to the coastal dunes of northern California J. Lep. Soc. 43 (3): 229-238
de Prunner, 1798
Lepidoptera Pedemontana illustrata. Turin. 1 Lepid. Pedemont. : 1-124
[NAB]; Pyle, 1981
Field Guide to North American Butterflies, National Audubon Society
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Rebel, 1903
Studien über die Lepidopterenfauna der Balkanländer Ann. nat. Mus. Wien 18 : 123-347, pl. 3
Rothschild, 1917
Supplemental notes to Mr. Charles Oberthür's Faune des Lépidoptères de la Barbarie, with lists of the specimens contained in the Tring Museum Novit. Zool. 24 (1): 61-120, (2): 325-373, 393-409
Rothschild, 1925
Critical List of the collection of Algerian Lepidoptera of the late Captain N. J. E. Holl. Novit. Zool. 32 : 195-229
Rottemburg, 1775
Unmertungen zu den Hufnagelifchen Tabellen der Schmetterlinge Der Naturforscher 6 : 1-34
Heyne, [1894]; Rühl, 1894; Rühl, 1895
Die palaearktischen Grossschmetterling und ihre Naturgeschichte Palaearkt. Grossschm. 1 : 1-857
de Sagarra, 1924
Noves formes de Lepidòpters ibèrics Butll. Inst. Catal. Hist. nat. (2) 4 (9) : 198-204
de Sagarra, 1930
Anotacions a la lepidopterologia Ibérica V (2). Formes noves de lepidòpters ibérics Butll. Inst. catal. Hist. nat. (2) 10 (7) : 110-118
Schwingenschuss, 1939
Beitrag zur Lepidopterenfauna von Iran (Persien), insbesondere des Elburs-Gebirges in Nordiran Ent. Zs. 53 (2): 13-14, (5): 36-38, (8): 62-64, (11): 86-88, (12): 95-96, (16): 125-128, (17): 135-136, (19): 149-152, (21): 166-168, (25): 200-210 (1940)
Seitz in Seitz, 1908; Seitz, 1908
Die Großschmetterlinge des Palaearktischen Faunengebietes. 1. Die Palaearktischen Tagfalter Gross-Schmett. Erde 1 : 1-379, pl. 1-89
Fruhstorfer in Seitz, 1911
Die Großschmetterlinge des Indo-australischen Faunengebietes Gross-Schmett. Erde 9 : 1-766, : 767-1197
[SPRK]; Seppänen, 1970
Suomen suurperhostoukkien ravintokasvit Animalia Fennica 14
Sheljuzhko, 1929
Einige neue palaearktische Lepidopteren-Formen Mitt. Münch. Ent. Ges. 19 : 347-362, pl. 27-28
Sheljuzhko, 1937
Lepidopterologische Ergebnisse meiner Reise nach dem Teberda-Gebiete (Nordwest-Kaukasus) Festschr. Embrik Strand, Riga 2 : 322-354
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Staudinger, 1861
Catalogue der Lepidopteren Europas und der angrendzenden Länder in Staudinger & Wocke, Cat. Lep. (ed. 1) : 1-192
Staudinger, 1870
Beschreibung neuer Lepidopteren des europäischen faunegebiets Berl. Ent. Z. 14 : 97-132, 193-208, (3/4): 273-330 (1871)
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Staudinger, 1892
Lepidopteren des Kentei-Gebirges Dt. Ent. Z. Iris 5 (2) : 300-393
Staudiger in Staudinger & Rebel, 1901; Staudinger in Staudinger & Rebel, 1901
Catalog der Lepidopteren des palaearktischen Faunengebiets. I. Famil. Papilionidae - Hepialidae Cat. Lep. palaearct. Faunengeb. 1 : 1-411
Staudinger in Staudinger & Wocke, 1871
Catalog der Lepidopteren des europaeischen Faunegebiets (Edn. 2): - Cat. Lepid. eur. Faunegeb. (Edn. 2) : 1-421, : 422-426 (Corrigenda et Addenda)
Stetter-Stättermayer, 1933
Ueber aberrative Formen von Coenonympha arcanioides Pier. Int. Ent. Zs. 27 (30) : 340
Stoll, [1782]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Strand, 1927
Neubenennungen palaearktischer Lepidoptera und Apidae Arch. Naturg. 91 A (12) : 281-283
Sugitani, 1932
Some new aberrant forms of Japanese and Formosan butterflies [in Japanese] Zephyrus 4 (1): 5-9, pl. 2, (?): 88-91
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
Thunberg, 1791
Dissertio Entomologica sistens Insecta Suecica, ... Diss. Ent. sistens Insecta Suecica (1): 1-24 (1 plate) (1784), (2): [25-31] ??], 32-46 (9 unnumbered) (1791), (3): [47],48-52 (2 unnumbered) (1792), (4): 53-62 (1792)
Tshikolovets, 2017
New taxa and new records of butterflies (Lepidoptera: Pieridae, Lycaenidae, Nymphalidae) from Afghanistan Zootaxa 4358 (1) : 107-124
Tutt, 1904
Coenonympha mathewi, n.sp. Ent. Rec. J. Var. 16 : 308-309
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Verity, 1914
Contributo allo studio della variazione nei Lepidotteri Boll. Soc. ent. ital. 45 : 203-238, pl. 1
Verity, 1916
The Brish Races of Butterflies: their relationships and nomeclature Ent. Rec. J. Var. 28 (4): 73-80, (5): 97-102, (6): 128-133, (7-8): 165-174
Wagner, 1913
Beitrag zur Lepidopterenfauna des Iligebietes sowie des Sary-Dschas (Asia centr.) Ent. Mitt. 2 (1): 22-30, (2): 50-62, (3): 88-95, (4): 112-126, (5): 153-158, (6): 185-190, (7/8): 244-254, (9): 285-288
Webster, 1998
The life history of the Maritime Ringlet, Coenonympha tullia nipisquit (Satyridae) J. Lep. Soc. 52 (4) : 345-355
Wyatt, 1952
Einige neue Tagfalterformen aus Marokko Zs. Wiener ent. Ges. 37 : 173-175
Wyatt, 1952
Einige neue Tagfalterrassen aus Spanien Zs. Wiender ent. Ges. 37 : 204-207
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96
Zhang, Cong, Shen, Opler & Grishin, 2020
Genomic evidence suggests further changes of butterfly names Taxonomic Rep. 8 (7) : 1-41

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