Spialia

Spialia Swinhoe, [1912] Grizzled Skippers, Sandmen

Spialia Swinhoe, [1912]; in Moore, Lepidoptera Indica 10: 99; TS: Hesperia galba Fabricius

Spialia; [LepIndex]; [CoL] ; [iNaturalist] ; [iNaturalist] ; [NHM-BMW]
Spialia (Pyrginae); [BK], 403; [AfrL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; [RFEL]

The asterodia species-group

Asterodia; de Jong, 1978, Tijdschr. Ent. 121: 32, 33

Spialia asterodia (Trimen, 1864)

Lesotho, Natal, Transvaal, Orange Free State, Cape, Mozambique, Rhodesia. See [maps]

Pyrgus asterodia Trimen, 1864; Trans. ent. Soc. Lond. (3) 2: 178; TL: "Cape Colony"; Plettenberg Bay

Spialia asterodia; de Jong, 1978, Tijdschr. Ent. 121: 33, pl. 1, f. 1-2; [PBSA]; [AfrL]; [LepIndex] ; [CoL]
Syrichtus asterodia; [BOW]: pl. 130, f. 42

Spialia agylla (Trimen, 1889)

-. See [maps]

Pyrgus agylla Trimen, 1889; South-African Butts 3: 286; TL: Griqualand West

Spialia agylla; de Jong, 1978, Tijdschr. Ent. 121: 36, pl. 1, f. 3-4; [PBSA]; [AfrL]; [LepIndex] ; [CoL]
Syrichtus agylla; [BOW]: pl. 130, f. 41

S. a. agylla South Africa, Botswana

Spialia agylla agylla; de Jong, 1978, Tijdschr. Ent. 121: 38; [AfrL]

S. a. bamptoni Vári, 1976 Namaqualand

Spialia agylla bamptoni Vári, 1976; Ann. Transv. Mus. 30 (13): 123, f. 3-4, pl. 13, f. 3-4; TL: Hondekipbaai

Spialia agylla bamptoni; de Jong, 1978, Tijdschr. Ent. 121: 38; [AfrL]; [LepIndex]; [CoL]

Spialia kituina (Karsch, 1896)

Kenya. See [maps] Kitui Grizzled Skipper

Hesperia kituina Karsch, 1896; Ent. Nachr. 22 (24): 374; TL: Kitui, Kenya
Pyrgus bettoni Butler, 1898; Proc. zool. Soc. Lond. 1898: 415, pl. 32, f. 1; TL: Maungu Inkubwa, Kenya
- = ; de Jong, 1978, Tijdschr. Ent. 121: 38

Spialia kituina; de Jong, 1978, Tijdschr. Ent. 121: 38, pl. f. 5-6; [BK]: 403, pl. 59, f. 766; [AfrL]; [LepIndex] ; [CoL]

The spio species-group

Spio; de Jong, 1978, Tijdschr. Ent. 121: 32, 41

Spialia spio (Linnaeus, 1764)

Tropical Africa, SW.Arabia. See [maps] Spio Grizzled Skipper

Papilio spio Linnaeus, 1764; Mus. Lud. Ulr.: 338
Papilio spio Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 796; TL: Cape of Good Hope
Papilio vindex Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 122, pl. 353, f. G, H; TL: Cape of Good Hope
- = ; de Jong, 1978, Tijdschr. Ent. 121: 41

Spialia spio; de Jong, 1978, Tijdschr. Ent. 121: 41, pl. 1, f. 7-8; [PBSA]; [BK]: 404, pl. 59, f. 767; [AfrL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
Syrichtus spio; [BOW]: pl. 130, f. 50

Spialia diomus (Hopffer, 1855)

-. See [maps]

Pyrgus diomus Hopffer, 1855; Ber. Verh. Akad. Berlin 1855: 643; TL: Mozambique

Spialia diomus; de Jong, 1978, Tijdschr. Ent. 121: 45, pl. 2, f. 9-12; [AfrL]; [LepIndex] ; [CoL] ; [iNaturalist]
Syrichtus diomus; [BOW]: pl. 130, f. 44

S. d. diomus W.Africa, Nigeria - Uganda, S.Sudan, Ethiopia, Somalia, Yemen, E.Africa Diomus Grizzled Skipper

Spialia diomus diomus; de Jong, 1978, Tijdschr. Ent. 121: 46, pl. 2, f. 9-10; [BK]: 404, pl. 59, f. 768; [AfrL]

S. d. ferax (Wallengren, 1863) South Africa, Botswana, Transvaal, Natal, Rhodesia, Moçambique

Syrichtus ferax Wallengren, 1863; Wien ent. Monats. 7 (5): 137; TL: Kuisip River
Pyrgus abscondita Plötz, 1884; Mitt. nat. Ver. Neu-Pomm. 15: 21; TL: Africa
- = ; de Jong, 1978, Tijdschr. Ent. 121: 49
Syrichtus lacreuzei Oberthür, 1912; Étud. Lépid. Comp. 6: 117, f. 1332; TL: Zoutpansberg, South Africa
- = ; de Jong, 1978, Tijdschr. Ent. 121: 49

Spialia abscondita; [LepIndex] ; [CoL]
Spialia diomus ferax; de Jong, 1978, Tijdschr. Ent. 121: 49, pl. 2, f. 11-12; [PBSA]; [AfrL]; [LepIndex]; [CoL]
Spialia ferax; [iNaturalist]
Syrichtus abscondita; [BOW]: pl. 130, f. 40

The phlomidis species-group

Phlomidis; de Jong, 1974, Tijdschr. Ent. 117: 231; de Jong, 1978, Tijdschr. Ent. 121: 32, 50

Spialia phlomidis (Herrich-Schäffer, [1845])

Balkans, S.Russia, Asia Minor, Iran. See [maps] Persian Skipper

Hesperia phlomidis Herrich-Schäffer, [1845]; Syst. Bearb. Schmett. Europ. 1 (11): 153, (12):(iii) pl. 2, f. 8-9; TL: (shores) Sea of Marmora, Turkey
jason Oberthür, 1912
eupator Hemming, 1932

Pyrgus phlomidis; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 502
Spialia phlomidis; de Jong, 1974, Tijdschr. Ent. 117: 233; [H&R]; de Jong, 1978, Tijdschr. Ent. 121: 52; [BOW]: pl. 11, f. 30; [BRU], 115; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE]

S. p. phlomidis Bulgaria, Greece, Turkey, S.Russia;Iran, Alai

Spialia phlomidis phlomidis; de Jong, 1974, Tijdschr. Ent. 117: 234; de Jong, 1978, Tijdschr. Ent. 121: 52

S. p. hermona Evans, 1956 Lebanon

phlomidis hermona Evans, 1956; Ann. Mag. nat. Hist. (12) 9 (106): 750; TL: Mt. Hermon
?Spialia phlomidis kiki Higgins; J. Ent. (B) 43 (1): (83-87)

Spialia phlomidis hermona; de Jong, 1974, Tijdschr. Ent. 117: 235; de Jong, 1978, Tijdschr. Ent. 121: 52; [LepIndex]; [CoL] ; [iNaturalist]
Spialia phlomidis kiki; [LepIndex]; [CoL]

Spialia osthelderi (Pfeiffer, 1932)

-. See [maps]

Hesperia (Ateleomorpha) osthelderi Pfeiffer, 1932; Mitt. Münch. Ent. Ges. 22: 50, pl. 4, f. 14-15, 18; TL: Marash, Taurus Mts, S. Turkey

Spialia osthelderi; de Jong, 1974, Tijdschr. Ent. 117: 235; de Jong, 1978, Tijdschr. Ent. 121: 52; [BRU], 115; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; [iNaturalist] ; [LEPIFORUM]
Spialia struvei osthelderi; [LepIndex]; [CoL]

S. o. osthelderi Asia Minor - Middle East, Syria, Lebanon, Iraq, W.Iran

Spialia osthelderi osthelderi; de Jong, 1974, Tijdschr. Ent. 117: 236

S. o. gecko Evans, 1949 Kopet-Dagh, Iran - Afghanistan, Sussamyr Mts

osthelderi gecko Evans, 1949; Cat. Hesp. Europe Asia Australia Brit. Mus.: 176; TL: Hyrcania
Spialia osthelder struveoides Brandt, 1939; (nom. nud., Pfeiffer)

Spialia geron gecko; [LepIndex]; [CoL]
Spialia osthelderi gecko; de Jong, 1974, Tijdschr. Ent. 117: 236; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10

Spialia geron (Watson, 1893)

Afghanistan, W.Pakistan. See [maps]

geron Watson, 1893; Proc. Zool. Soc. Lond. 1893 (1): 66; TL: Quetta, Baluchistan

Spialia geron geron; de Jong, 1974, Tijdschr. Ent. 117: 238
Spialia geron; de Jong, 1974, Tijdschr. Ent. 117: 238; de Jong, 1978, Tijdschr. Ent. 121: 52; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; [LepIndex] ; [CoL] ; [LEPIFORUM]

Spialia struvei (Püngeler, 1914)

Iran - Dzhungaria. See [maps]

Hesperia struvei Püngeler, 1914; Dt. ent. Z. Iris 28 (1): 37, pl. 2, f. 13, 20; TL: Barkul, E. Tian-Shan

Spialia geron struvei; de Jong, 1974, Tijdschr. Ent. 117: 238; de Jong, 1978, Tijdschr. Ent. 121: 53; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10
Spialia struvei; [BRU], 115; Yakovlev, 2012, Nota lepid. 35 (1): 60; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

S. s. irida Zhdanko, 1993 Alaisky Mts.

Spialia geron irida; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10
Spialia irida; [LEPIFORUM]

S. s. fetida Zhdanko, 1992 W.Kazakhstan

?Spialia fetida Zhdanko, 1992; Trudy Zool. inst. 248: (184-186)

Spialia fetida; [LEPIFORUM]
Spialia geron fetida; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10

Spialia doris (Walker, 1870)

Arabia, N.India, E.Africa. See [maps] Desert Grizzled Skipper Aden Skipper

Nisoniades doris Walker, 1870; Entomologist 5 (76): 56; TL: Tajora (Straits of Bab-el-Mendeb) [Egypt]

Spialia doris; de Jong, 1974, Tijdschr. Ent. 117: 239; de Jong, 1978, Tijdschr. Ent. 121: 53; [H&R], 325; [BRU], 116; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; [AfrL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

S. d. doris NE.Kenya, S.Arabia, Somalia, Ethiopia

Spialia evanidus adenensis Butler, 1884; Proc. zool. Soc. Lond. 1884: 493; TL: Aden

Spialia doris doris; de Jong, 1974, Tijdschr. Ent. 117: 241; de Jong, 1978, Tijdschr. Ent. 121: 53 (note); [BK]: 404, pl. 59, f. 769; [AfrL]; [iNaturalist]

S. d. daphne Evans, 1949 SW.Morocco

Spialia doris daphne Evans, 1949; Cat. Hesp. Europe Asia Australia Brit. Mus.: 177; TL: High Atlas, Ziz Valley (Morocco)

Spialia doris daphne; de Jong, 1974, Tijdschr. Ent. 117: 240; [H&R], 325; [BMAT]: 92, pl. 30, f. 34-37; [AfrL]; [LepIndex]; [CoL] ; [iNaturalist]

S. d. amenophis (Reverdin, 1914) N.Egypt

doris amenophis (Reverdin, 1914); Bull. Soc. Lép. Genève 3: 55, pl. 3, f. 5, 11, pl. 4, f. 1; TL: Heliopolis, Egypt

Spialia doris amenophis; de Jong, 1978, Tijdschr. Ent. 121: 54 (note); [BMAT]: 93 (note); [AfrL]; [LepIndex]; [CoL]

S. d. evanida (Butler, 1880) S.Turkmenia, S.Uzbekistan, S.Iran

doris evanida Butler, 1880; Ann. Mag. nat. Hist. (5) 5 (27): 223; TL: Hubb River, S. Baluchistan

Spialia doris evanida; de Jong, 1974, Tijdschr. Ent. 117: 241; de Jong, 1978, Tijdschr. Ent. 121: 54 (note); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10

The colotes species-group

Colotes; de Jong, 1978, Tijdschr. Ent. 121: 32, 54

Spialia colotes (Druce, 1875)

-. See [maps]

Pyrgus colotes Druce, 1875; Proc. zool. Soc. Lond. 1875: 416; TL: Angola

Spialia colotes; de Jong, 1978, Tijdschr. Ent. 121: 54, pl. 2, f. 13-14; [BOW]: pl. 130, f. 38; [AfrL]; [LepIndex] ; [CoL]

S. c. colotes Angola

Pyrgus nora Plötz, 1884; Mitt. nat. Ver. Neu-Pomm. 15: 7; TL: Loango, Angola
- = ; de Jong, 1978, Tijdschr. Ent. 121: 55

Spailia colotes colotes; de Jong, 1978, Tijdschr. Ent. 121: 55
Spialia colotes colotes; [AfrL]

S. c. transvaaliae (Trimen, 1889) Botswana, Transvaal, Rhodesia - Kenya, Uganda Transvaal Grizzled Skipper

Pyrgus transvaaliae Trimen, 1889; South-African Butts 3: 286; TL: Transvaal

Spialia colotes transvaaliae; de Jong, 1978, Tijdschr. Ent. 121: 56; [BK]: 405, pl. 59, f. 770; [AfrL]; [LepIndex]; [CoL]

S. c. semiconfluens de Jong, 1978 Ethiopia, Somalia, SW.Arabia

Spialia colotes semiconfluens de Jong, 1978; Tijdschr. Ent. 121: 58, pl. 2, f. 13-14; TL: Bihendula, Somalia

Spialia colotes semiconfluens; [BK], 405 (note); [AfrL]; [LepIndex]; [CoL]

Spialia confusa (Higgins, 1924)

-. See [maps]

Hesperia transvaaliae var. confusa Higgins, 1924; Trans. ent. Soc. Lond. 1924: 90; TL: SW shore Lake Nyase, betw. Ft. Johnston and Monkey Bay

Spialia confusa; de Jong, 1978, Tijdschr. Ent. 121: 59, pl. 2, f. 15-16; [AfrL]; [LepIndex] ; [CoL]

S. c. confusa Tanzania - Natal, N.Zambia

Hesperia transvaaliae var. fasciata Higgins, 1924; Trans. ent. Soc. Lond. 1924: 90; TL: betw. Ft. Mangoche and Chikala Boma, Malawi
- = ; de Jong, 1978, Tijdschr. Ent. 121: 60

Spialia confusa confusa; de Jong, 1978, Tijdschr. Ent. 121: 60; [AfrL]

S. c. obscura (Higgins, 1924) Kenya

Hesperia transvaalieae var. obscura Higgins, 1924; Trans. ent. Soc. Lond. 1924: 92; TL: Nairobi

Spialia confusa obscura; de Jong, 1978, Tijdschr. Ent. 121: 59, pl. 2, f. 15-16; [BK]: 405, pl. 59, f. 771; [AfrL]; [LepIndex]; [CoL]

The sertorius species-group

Sertorius; de Jong, 1974, Tijdschr. Ent. 117: 241; de Jong, 1978, Tijdschr. Ent. 121: 32, 61

Spialia mafa (Trimen, 1870)

Natal, Botswana, Rhodesia, South West Africa, Cape, S.Africa - Sudan, S.Arabia. See [maps]

Pyrgus mafa Trimen, 1870; Trans. ent. Soc. Lond. 1870 (4): 386, pl. 6, f. 12; TL: Maseru, Basutoland

Spialia mafa; de Jong, 1978, Tijdschr. Ent. 121: 62, pl. 4, f. 25-26; [PBSA]; [AfrL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

S. m. mafa S.Zaire, Zambia, Malawi, S.Africa

Hesperia oberthüri Aurivillius, 1925; in Seitz, Gross-Schmett. Erde 13: 565; TL: Kimberley
- = ; de Jong, 1978, Tijdschr. Ent. 121: 65
Pyrgus aurivillii Shepard, 1935; Lepid. Cat. 69: 496 (repl. Hesperia oberthüri Aurivillius, 1925)

Spialia mafa mafa; de Jong, 1978, Tijdschr. Ent. 121: 65; [AfrL]; [iNaturalist]

S. m. higginsi Evans, 1937 Oman, Yemen, Ethiopia, Kenya, Uganda, Tanzania, Malawi Mafa Grizzled Skipper

Spialia mafa higginsi Evans, 1937; Cat. African Hesp. Brit. Mus.: 62; TL: Mumia, Kenya

Spialia mafa higginsi; de Jong, 1978, Tijdschr. Ent. 121: 64; [BK]: 405, pl. 59, f. 772; [AfrL]; [LepIndex]; [CoL] ; [iNaturalist]

Spialia galba (Fabricius, 1793)

-. See [maps] Indian Grizzled Skipper Indian Skipper

Hesperia galba Fabricius, 1793; Ent. Syst. 3 (1): 352; TL: Tranquebar, S.India

Hesperia galba; Moore, [1881], Lepid. Ceylon 1 (4): 183, pl. 71, f. 6
Spialia galba; de Jong, 1978, Tijdschr. Ent. 121: 66, pl. 4, f. 27-28; [BOW]: pl. 188, f. 41; [MRS], 716; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
Syrichtus galba; [BIR], 466

732x539 (~49Kb) Thailand, Chon Buri Province, Pattaya, Ko Khram Island, Saikawe beach, a side of a road through shrubbery at the coast, 10th January 2006, Photo © Oleg Kosterin

794x624 (~51Kb) Thailand, Chon Buri Province, Pattaya, Ko Khram Island, Saikawe beach, a side of a road through shrubbery at the coast, 10th January 2006, Photo © Oleg Kosterin

S. g. galba Pakistan, India, Kashmir, Nepal, Sikkim, Bhutan, Ceylon, Thailand

Pyrgus superna Moore, [1866]; Proc. zool. Soc. Lond. 1865 (3): 792; TL: Calcutta
- = ; de Jong, 1978, Tijdschr. Ent. 121: 68

Spialia galba galba; de Jong, 1978, Tijdschr. Ent. 121: 68; [iNaturalist]

S. g. shanta Evans, 1956 Burma

Spialia galba shanta Evans, 1956; Ann. Mag. nat. Hist. (12) 9 (106): 750; TL: Hsipaw, S. Shan States

Spialia galba shanta; de Jong, 1978, Tijdschr. Ent. 121: 69; [LepIndex]; [CoL] ; [Yutaka Inayoshi]

S. g. chenga Evans, 1956 Hainan

Spialia galba chenga Evans, 1956; Ann. Mag. nat. Hist. (12) 9 (106): 750; TL: Kiung-Chow, Hainan

Spialia galba chenga; de Jong, 1978, Tijdschr. Ent. 121: 69; [LepIndex]; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]

Spialia sertorius (Hoffmannsegg, 1804)

W.Asia?, Chitral?, Altai?, Tibet?, Amur?. See [maps] Red Underwing Skipper Sertorio Powszelatek sertor

Hesperia sertorius Hoffmannsegg, 1804; Mag. f. Insektenk. 3: 203; TL: Germany

Spialia sertorius; de Jong, 1974, Tijdschr. Ent. 117: 246; de Jong, 1978, Tijdschr. Ent. 121: 71; [H&R], 324; [BOW]: pl. 11, f. 31-32; [BMAT]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10 (?); [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE]

366x348 (~39Kb) Mt. Canigou, Pyrenees Orientales, France 27/7/96, Photo © Simon Coombes

450x363 (~35Kb) upperside Mt. Canigou, Pyrenees Orientales, France 27/7/96, Photo © Simon Coombes

S. s. sertorius CEU, SEU

Papilio sao Hübner, ?; (invalid homonym)
- = ; de Jong, 1974, Tijdschr. Ent. 117: 250
hibiscae Hemming, 1936
- = ; de Jong, 1974, Tijdschr. Ent. 117: 250
sertorius parvula Verity
- = ; de Jong, 1974, Tijdschr. Ent. 117: 250
hibiscae minor Rebel
- = ; de Jong, 1974, Tijdschr. Ent. 117: 250
gracilis Verity, 1921; ; TL: Florence
- = ; de Jong, 1974, Tijdschr. Ent. 117: 250
subgracilis Verity, 1912
- = ; de Jong, 1974, Tijdschr. Ent. 117: 250
guadarramensis Warren, 1925; ; TL: La Granja, Sierra de Guadarrama
- = ; de Jong, 1974, Tijdschr. Ent. 117: 251
gavarniensis Warren, 1926; ; TL: Gavarnie
- = ; de Jong, 1974, Tijdschr. Ent. 117: 251
alioides Verity, 1926; ; TL: Oulx, Cottian Alps
- = ; de Jong, 1974, Tijdschr. Ent. 117: 251

Spialia sertorius sertorius; de Jong, 1974, Tijdschr. Ent. 117: 248; [H&R], 324

S. s. ali (Oberthür, 1881) Morocco, Algeria, Tunisia

Syrichtus ali Oberthür, 1881; Étud. d'Ent. 6: 61, pl. 2, f. 3; TL: Oran Constantine [Lambessa] (Algeria)
Syrichtus sao therapnoïdes Oberthür, 1910; Ètud. Lépid. Comp. 4: 385, pl. 54, f. 448; TL: Sebdou (Algeria)
Spialia sertorius ali f. rungsi Picard, 1950; Bull. Soc. Sci. nat. Maroc 28: 123; TL: Ifrane (Morocco)

Spialia ali; [iNaturalist] ; [LEPIFORUM]
Spialia sertorius ali; de Jong, 1974, Tijdschr. Ent. 117: 247; [H&R], 325; [BMAT]: 92, pl. 30, f. 29-33; [LepIndex]; [CoL]

Spialia therapne (Rambur, 1832)

Corsica, Sardinia. See [maps]

Hesperia therapne Rambur, 1832; Ann. Soc. ent. Fr. 1: 265, pl. 7, f. 4; TL: Corsica

Spialia sertorius therapne; de Jong, 1974, Tijdschr. Ent. 117: 248; [H&R], 324; [LepIndex]; [CoL]
Spialia therapne; [Otakar Kudrna]; [iNaturalist] ; [LEPIFORUM] ; [MBE]

Spialia orbifer (Hübner, [1823])

Temperate Asia - Korea. See [maps]

Hesperia orbifer Hübner, [1823]; Samml. eur. Schmett. [1]: pl. 161, f. 803-806; TL: Hungary
?Hesperia orbifer murasaki Sugitani, 1936; Zephyrus 6: 155

Spialia orbifer; de Jong, 1974, Tijdschr. Ent. 117: 252; de Jong, 1978, Tijdschr. Ent. 121: 71; [BRU], 116; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; [RFEL]; [iNaturalist] ; [LEPIFORUM] ; [MBE]

1296x1039 (~275Kb) ♂ upperside Two males on a fresh horse dung. A path in an open larch forest in the forest belt upper part, south-estern slope of the Tanmalyg River valley, Khan-Deer cluster of the Ubsunurskaya Kotlovina State Biosphaera Nature Reserve, the Khor-Taiga Mts. within the Khemchik Mountain Range, West Sayan, Chaa-Khol Kozhuun of Tyva Republic (Tuva), South Siberia, Russia. 4th July 2004, Photo © Oleg Kosterin

560x617 (~93Kb) ♂ underside A male on a fresh horse dung. A path in an open larch forest in the forest belt upper part, south-estern slope of the Tanmalyg River valley, Khan-Deer cluster of the Ubsunurskaya Kotlovina State Biosphaera Nature Reserve, the Khor-Taiga Mts. within the Khemchik Mountain Range, West Sayan, Chaa-Khol Kozhuun of Tyva Republic (Tuva), South Siberia, Russia. 4th July 2004, Photo © Oleg Kosterin

S. o. orbifer SEEU - SW.Siberia, N.Iran, Sicily

tesselloides Herrich-Schäffer, [1845]; ; TL: Sicily
- = ; de Jong, 1974, Tijdschr. Ent. 117: 254
f. minor Rebel, 1909; (gen. aest.)
- = ; de Jong, 1974, Tijdschr. Ent. 117: 253
posttesselloides Verity, 1938; ; TL: Olympus
- = ; de Jong, 1974, Tijdschr. Ent. 117: 253

Spialia orbifer orbifer; de Jong, 1974, Tijdschr. Ent. 117: 253; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10
Spialia sertorius orbifer; [H&R], 324; [LepIndex]; [CoL]

S. o. hilaris (Staudinger, 1901) SE.Turkey - Lebanon, Israel, Jordan, N.Iraq, W.Iran

orbifer hilaris Staudinger, 1901; Cat. Lepid. palaearct. Faunengeb.: 96; TL: N.Mesopotamia
secunda Graves, 1925
- = ; de Jong, 1974, Tijdschr. Ent. 117: 255

Spialia hilaris; [LEPIFORUM]
Spialia orbifer hilaris; de Jong, 1974, Tijdschr. Ent. 117: 254; [iNaturalist]
Spialia sertorius hilaris; [LepIndex]; [CoL]

S. o. carnea (Reverdin, 1927) Afghanistan, Baluchistan, Chitral

orbifer carnea Reverdin, 1927; Horae Macrolep. Palaearct. 1: 52, pl. 7, f. 28; TL: Paghman Mountains, Afghanistan

Spialia carnea; [LEPIFORUM]
Spialia orbifer carnea; de Jong, 1974, Tijdschr. Ent. 117: 256; de Jong, 1978, Tijdschr. Ent. 121: 71; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10
Spialia sertorius carnea; [LepIndex]; [CoL]

S. o. lugens (Staudinger, 1886) Tian-Shan, NE.Iran, Transcaspia, S.Siberia - Amur

Pyrgus orbifer var. lugens Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 256; TL: Ferghana, E.Tian-Shan

Pyrgus orbifer var. lugens; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 505
Spialia lugens; ; [LEPIFORUM]
Spialia orbifer lugens; de Jong, 1974, Tijdschr. Ent. 117: 255; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10
Spialia sertorius lugens; [LepIndex]; [CoL]

S. o. pseudolugens Gorbunov, 1995 Altai, S.Urals

orbifer pseudolugens Gorbunov, 1995; ; TL: Altai and S.Urals

Spialia orbifer pseudolugens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 10; Yakovlev, 2012, Nota lepid. 35 (1): 60; [RFEL]

The dromus species-group

Dromus; de Jong, 1978, Tijdschr. Ent. 121: 32, 72

Spialia wrefordi Evans, 1951

N.Kenya, Uganda. See [maps] Wreford's Grizzled Skipper

Spialia wrefordi Evans, 1951; Ann. Mag. nat. Hist. (12) 4: 1270; TL: Kotido, Karamoja, Uganda

Spialia wrefordi; de Jong, 1978, Tijdschr. Ent. 121: 73, pl. 5, f. 33-34; [BK]: 405, pl. 59, f. 773; [AfrL]

Spialia paula (Higgins, 1924)

Botswana, Transvaal, Rhodesia. See [maps]

Hesperia paula Higgins, 1924; Trans. ent. Soc. Lond. 1924: 77, pl. 8, f. 22; TL: Bulawayo, Rhodesia

Spialia paula; de Jong, 1978, Tijdschr. Ent. 121: 74, pl. 5, f. 35-36; [AfrL]; [LepIndex] ; [CoL]

Spialia secessus (Trimen, 1891)

Tanzania, Rhodesia, Transvaal. See [maps]

Pyrgus secessus Trimen, 1891; Proc. zool. soc. Lond. 1891: 102 [dry-season], pl. 9, f. 22; TL: Omrora, Damara-land
Hesperia secessa f. trimeni Aurivillius, 1925; in Seitz, Gross-Schmett. Erde 13: 564, pl. 75 d [wet-season]; TL: Rhodesia

Spialia secessus; de Jong, 1978, Tijdschr. Ent. 121: 76, pl. 5, f. 37-38; [AfrL]; [LepIndex] ; [CoL]
Syrichtus secessus; [BOW]: pl. 130, f. 49

Spialia dromus (Plötz, 1884)

Transkei - Natal (coastal), Tropical Africa. See [maps] Dromus Grizzled Skipper

Pyrgus dromus Plötz, 1884; Mitt. nat. Ver. Neu-Pomm. 15: 6; TL: Congo
Pyrgus zaira Plötz, 1884; Mitt. nat. Ver. Neu-Pomm. 15: 6; TL: Congto
- = ; de Jong, 1978, Tijdschr. Ent. 121: 80
Syrichthus melaleuca Oberthür, 1912; Étud. Lépid. Comp. 6: 113, f. 1327; TL: Kitang
- = ; de Jong, 1978, Tijdschr. Ent. 121: 80
Syrichthus leucomelas Oberthür, 1912; Étud. Lépid. Comp. 6: 118, f. 1308; TL: Mpala, L. Tanganyika
- = ; de Jong, 1978, Tijdschr. Ent. 121: 80
Hesperia dromus var. elongata Higgins, 1924; Trans. ent. Soc. Lond. 1924: 95; TL: Makindu, Kenya
- = ; de Jong, 1978, Tijdschr. Ent. 121: 80
Hesperia dromus var. meridionalis Higgins, 1924; Trans. ent. Soc. Lond. 1924: 95; TL: Zanzibar
- = ; de Jong, 1978, Tijdschr. Ent. 121: 80

Spialia dromus; de Jong, 1978, Tijdschr. Ent. 121: 80, pl. 5, f. 39-40; [PBSA]; [BOW]: pl. 130, f. 39; [BK]: 406, pl. 59, f. 774; [AfrL]; [LepIndex] ; [CoL]

Spialia ploetzi (Aurivillius, 1891)

W.Kenya - W.Africa. See [maps]

Hesperia ploetzi Aurivillius, 1891; Ent. Tidskr. 12: 227; TL: Cameroun

Spialia ploetzi; de Jong, 1978, Tijdschr. Ent. 121: 84; [AfrL]; [LepIndex] ; [CoL] ; [LEPIFORUM]
Syrichtus ploetzi; [BOW]: pl. 130, f. 46

S. p. ploetzi W.Cameroun, W.Kenya, Uganda, Burundi, Rwanda, Zaire Forest Grizzled Skipper

Hesperia zebra Rebel, 1914; Ann. Mus. Wien. 28: 271, pl. 22, f. 52-53 (preocc. Pyrgus zebra Butler, 188)
Hesperia rebeli Higgins, 1924; Trans. ent. Soc. Lond. 1924: 101 (repl. Hesperia zebra Rebel, 1914)

Spialia ploetzi ploetzi; de Jong, 1978, Tijdschr. Ent. 121: 85, pl. 6, f. 41-42; [BK]: 406, pl. 59, f. 775; [AfrL]
Syrichtus rebeli; [BOW]: pl. 130, f. 47

S. p. occidentalis de Jong, 1977 W.Africa

Spialia ploetzi occidentalis de Jong, 1977; Ent. Ber. Amst. 37: 43; TL: Ghana

Spialia ploetzi occidentalis; de Jong, 1978, Tijdschr. Ent. 121: 87; [BK]: 406 (note); [AfrL]; [LepIndex]; [CoL]

The delagoae species-group

Delagoae; de Jong, 1978, Tijdschr. Ent. 121: 32, 88

Spialia mangana (Rebel, 1899)

SW.Arabia, Somalia, Ethiopia, N.Kenya, Uganda (Karamoja). See [maps] Arabian Grizzled Skipper

Hesperia (Pyrgus) mangana Rebel, 1899; Anz. Akad. Wiss. Wien 36: 360; TL: Ras Fartak, S. Yemen

Spialia mangana; de Jong, 1978, Tijdschr. Ent. 121: 89, pl. 6, f. 45-46; [BK]: 406, pl. 59, f. 776; [AfrL]; [LepIndex] ; [CoL]

Spialia nanus (Trimen, 1889)

Cape, W.Orange Free State. See [maps]

Pyrgus nanus Trimen, 1889; South-African Butts 3: 290; TL: Malmesbury, Cape

Spialia nanus; de Jong, 1978, Tijdschr. Ent. 121: 91, pl. 6, f. 47-48; [PBSA]; [AfrL]; [LepIndex] ; [CoL]
Syrichtus nanus; [BOW]: pl. 130, f. 45

Spialia delagoae (Trimen, 1898)

South West Africa, Botswana, Transvaal, Swaziland, Zululand, Rhodesia, Moçambique, Kenya. See [maps] Delagoa Grizzled Skipper

Pyrgus delagoae Trimen, 1898; Trans. ent. Soc. Lond. 1898 (1): 15; TL: Delagoa Bay

Spialia delagoae; de Jong, 1978, Tijdschr. Ent. 121: 94, pl. 7, f. 49-50; [BK]: 406, pl. 59, f. 777; [AfrL]; [LepIndex] ; [CoL] ; [LEPIFORUM]
Syrichtus delagoae; [BOW]: pl. 130, f. 43

Spialia zebra (Butler, 1888)

-. See [maps]

Pyrgus zebra Butler, 1888; Ann. Mag. nat. Hist. (6) 1 (3): 207; TL: Koteer, Chittar Pahar

Spialia zebra; de Jong, 1978, Tijdschr. Ent. 121: 95, pl. 7, f. 51-52; [AfrL]; [LepIndex] ; [CoL]

S. z. zebra Pakistan, Punjab

Hesperia hellas de Nicéville, 1889; J. Bombay nat. Hist. Soc. 4 (3): 177, pl. B, f. 9; TL: Campbellpur (Punjab)
- = ; de Jong, 1978, Tijdschr. Ent. 121: 98

Spialia zebra zebra; de Jong, 1978, Tijdschr. Ent. 121: 98; [BK], 407 (note)

S. z. bifida (Higgins, 1924) Kenya, Uganda, Ethiopia, Somalia, SW.Arabia

Hesperia bifida Higgins, 1924; Trans. ent. Soc. Lond. 1924: 111; TL: Nairobi

Spialia zebra bifida; de Jong, 1978, Tijdschr. Ent. 121: 98; [BK]: 407, pl. 59, f. 778; [AfrL]; [LepIndex]; [CoL]

Spialia sataspes (Trimen, 1864)

Cape (Coastal). See [maps]

Pyrgus sataspes Trimen, 1864; Trans. ent. Soc. Lond. (3) 2: 178; TL: Plettenberg, Cape Prov.

Spialia sataspes; de Jong, 1978, Tijdschr. Ent. 121: 99, pl. 7, f. 53-54; [PBSA]; [AfrL]; [LepIndex] ; [CoL]
Syrichtus sataspes; [BOW]: pl. 130, f. 48

Spialia depauperata (Strand, 1911)

-. See [maps]

Hesperia ferax var. depauperata Strand, 1911; Mitt. zool. Mus. Berl. 5 (2): 304; TL: Msamwia, betw. Lake Rukwa and Lake Tanganyika, S.Tanzania

Spialia depauperata; de Jong, 1978, Tijdschr. Ent. 121: 102, pl. 7, f. 55-56; [AfrL]; [LepIndex] ; [CoL]

S. d. depauperata Uganda, Kenya - Zambia, Mozambique Deprived Grizzled Skipper

Syrichthus rehfousi Oberthür, 1912; Étud. Lépid. Comp. 6: 117, f. 1331; TL: Mpala, W. shore of Lake Tanganyika
- = ; de Jong, 1978, Tijdschr. Ent. 121: 104

Spialia depauperata depauperata; de Jong, 1978, Tijdschr. Ent. 121: 104; [BK]: 407, pl. 59, f. 779; [AfrL]

S. d. australis de Jong, 1978 Natal, Transvaal - Rhodesia

Spialia depauperata australis de Jong, 1978; Tijdschr. Ent. 121: 105; TL: Muden, Natal

Spialia depauperata australis; [AfrL]; [LepIndex]; [CoL]

References:

[AJA]
African Journal Archive
[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BHL] The
Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[EVERSMANNIA]
Eversmannia contents
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[SZMN] Lepidoptera
Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Aurivillius, 1891
Verzeichniss einer vom Herrn Fritz Theorin aus Gabun und dem Gebiete des Camerunflusses heimgebrachten Schmetterlings-sammlung Ent. Tidskr. 12 : 193-228, pl. 1-3
Aurivillius, 1925
Die Gross-Schmetteringe der Erde; 13 Band; Die Afrikanischen Tagfalter in Seitz, Gross-Schmett. Erde 13 : 1-613, pl. 1-80
Reverdin in Bang-Haas, 1927
Horae Macrolepidopterologicae Regionis Palaearcticae Horae Macrolep. Palaearct. 1 : 1-128, pl. 1-11
Butler, 1880
On a small collection of Lepidoptera from western India and Belochistan Ann. Mag. nat. Hist. (5) 5 (27) : 221-226
Butler, 1884
On a collection of Lepidoptera made by Major J.W. Yerbury at or near Aden Proc. zool. Soc. Lond. 1884 : 478-503, pl. 46
Butler, 1888
An Acccount of three Series of Lepidoptera collected in North-west India by Major Yerbury Ann. Mag. nat. Hist. (6) 1 (2): 132-151, (3): 196-209
Butler, 1898
On a collection of Lepidoptera made in British East Africa by Mr C. S. Betton Proc. zool. Soc. Lond. 1898 : 395-444, pl. 32-33
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
[PBSA]; Dickson & Kroon, 1978
Pennington's Butterflies of Southern Africa
Druce, 1875
A list of the Collection of Diurnal Lepidoptera made by J.J. Monteiro, Esq., in Angola, with Descriptions of some new Species Proc. zool. Soc. Lond. 1875 : 406-417
Evans, 1937
A catalogue of the African Hesperiidae. British Museum Cat. African Hesp. Brit. Mus. : 1-212, pl. 1-30
Evans, 1949
A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum Cat. Hesp. Europe Asia Australia Brit. Mus. : 1-502, pl. 1-53
Evans, 1951
Revisional notes on African Hesperiidae Ann. Mag. nat. Hist. (12) 4 : 1268-1272
Evans, 1956
Revisional Notes on the Hesperiidae of Europe, Asia and Australia Ann. Mag. nat. Hist. (12) 9 (106) : 749-752
Fabricius, 1793
Entomologia Systematica emendata et aucta. Ent. Syst. 3 (1) : 1-487
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Herrich-Schäffer, [1845]
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 (9): Title ("1843"), (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843), (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843), (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844), (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844), (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844), (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844), (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844), (9): 115-122 (1844), (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845), (11): 147-162 (1845), (12): (iii) pl. 1-3, f. 1-17 (1845), (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846), (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846), (26): (ii) pl. 71-77, f. 339-376 (1847), (32): (ii) pl. 78-82, f. 377-400 (1848), (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848), (37): (ii) pl. 90-91, f. 427-437 (1849), (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850), (45): (ii) pl. 95-102, f. 453-489 (1850), (47): (ii) pl. 103, f. 490-495 (1851), (49): (ii) pl. 104-109, f. 496-528 (1851), (51): (ii) pl. 110-118, f. 529-570 (1851), (54): (ii) pl. 119-123, f. 571-596 (1852), (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852), (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853), (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854), (66): (index) 21-24 (1855)
Higgins, 1924
On the Ethiopian species of the genus Hesperia (Lep. Rhop.), with descriptions of two new species Trans. ent. Soc. Lond. 1924: 64-120
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
Hoffmannsegg, 1804
Alphabetisches Verzeichniss zu J. Hübner's Abbildungen der Papilionen mit den beigefügten vorzüglichsten Synonymen Mag. f. Insektenk. 3 : 181-206
Hopffer, 1855
[Lepidoptera Diurna] Ber. Verh. Akad. Berlin 1855 : 639-643
Hübner, [1823]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 982-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
de Jong, 1974
Systematics and evolution of palaearctic Spialia species Tijdschr. Ent. 117 : 225-271
de Jong, 1977
Problems about Spiala ploetzi (Aurivillius) (Lepidoptera, Hesperiidae) Ent. Ber. Amst. 37 : 41-44
de Jong, 1978
Monograph of the genus Spialia Swinhoe Tijdschr. Ent. 121 : 23-146, pl. 1-7
Karsch, 1896
Aethiepische Hesperiiden Ent. Nachr. 22 (24) : 372-378
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1764
Museum S'ae R'ae M'tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque Mus. Lud. Ulr. : 1-720
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2) : 533-1327, : Errata
Moore, [1866]
On the Lepidopterous Insects of Bengal Proc. zool. Soc. Lond. 1865 (3) : 755-823, pl. 41-43
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Oberthür, 1910
Notes pour servir à établir la Faune Française et Algérienne des Lépidoptères Ètud. Lépid. Comp. 4 : 15-691, pl. 35-58
Oberthür, 1912
Observations sur les Hesperiidae du genre Syrichthus Étud. Lépid. Comp. 6 : 47-120, pl. 137-143
Pfeiffer in Osthelder & Pfeiffer, 1932
Lepidopteren-Fauna von Marasch in türkisch Nordsyrien Mitt. Münch. Ent. Ges. 22 : 17-32, pl. 3-4, : 38-51, : 52-73, : 74, : 74-82, : 83-87
Plötz, 1884
Analytische Tabellen der Hesperiinen-Gattungen Pyrgus und Carcharodus Mitt. nat. Ver. Neu-Pomm. 15 : 1-24
Püngeler, 1914
Neue palaearktische Macrolepidopteren Dt. ent. Z. Iris 28 (1) : 37-55, pl. 2-3
Rambur, 1832
Catalogue des Lépidoptéres de l'ile Corse, avec la description et la figure des espéces inédites (1) Ann. Soc. ent. Fr. 1 : 245-295
Rebel, 1899
Diagnosen neuer Lepidopteren aus Südarabien und von der Insel Sokotra Anz. Akad. Wiss. Wien 36 : 359-361
Rebel, 1914
Wissenschaftliche Ergebnisse der Expedition R. Grauernach. Zentralafrika. 1909-1911. Lepidoptera Ann. Mus. Wien. 28 : 219-294, pl. 17-24
Reverdin, 1914
Notes sur les genres Carcharodus, Hesperia et Thanaos Bull. Soc. Lép. Genève 3 : 38-77
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Strand, 1911
Zoologische Ergebnisse der Expedition des Herrn Hauptmann a. D. Fromm 1908/09 nach Deutsch-Ostafrica. I. Lepidoptera Mitt. zool. Mus. Berl. 5 (2) : 275-304
Sugitani, 1936
Corean butterflies (3[?]) Zephyrus 6 : 155-174
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
Trimen, 1864
Descriptions of some New Species of Butterflies found in Southern Africa Trans. ent. Soc. Lond. (3) 2 : 175-180
Trimen, 1870
Notes on Butterflies collected by J. H. Bowker, Esq., in Basuto-land, South Africa; with descriptions of some New Species Trans. ent. Soc. Lond. 1870 (4) : 341-390, pl. 6
Trimen, 1889
South-African Butterflies: a monograph of the extra-tropical species. Vol. 3 South-African Butts 3 : 1-438, pl. 10-12
Trimen, 1891
On Butterflies collected in tropical South-Western Africa, by Mr. A. W. Eriksson Proc. zool. soc. Lond. 1891 : 59-107, pl. 8-9
Trimen, 1898
On some new or little-known Species of African Butterflies Trans. ent. Soc. Lond. 1898 (1) : 1-16, pl. 1
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Vári, 1976
South African Lepidoptera 7. Descriptions and notes of new taxa of Rhopalocera Ann. Transv. Mus. 30 (13) : 121-144, pl. 13-17
Walker, 1870
A list of the Butterflies collected by J.K. Lord Esq. in Egypt, along the African shore of the Red Sea, and in Arabia; with descriptions of the Species new to Science Entomologist 5 (76): 48-57, (80-81): 123-134, (82): 151-155
Wallengren, 1863
Lepidopterologische Mittheilungen (1-3) Wien ent. Monats. 4 (2): 33-46, 4 (6): 161-176 (1860), 7 (5): 137-151 (1863)
Watson, 1893
A proposed classification of the Hesperiidae, with a revision of the genera Proc. Zool. Soc. Lond. 1893 (1) : 3-133, pl. 1-3
[BIR]; Wynter-Blyth, 1957
Butterflies of the Indian Region (1982 Reprint)
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96
Zhdanko, 1992
[A new skipper species of the genus Spialia Swinh. (Lepidoptera, Hesperiidae) from Kazakhstan] Trudy Zool. inst. 248: 184-186 (Russian)

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