Miletus

Miletus Hübner, [1819] Brownies

Miletus Hübner, [1819]; Verz. bek. Schmett. (5): 71; TS: Papilio symethus Cramer
Symetha Horsfield, [1828]; Descr. Cat. lep. Ins. Mus. East India Coy (1): pl. [2]; TS: Symetha pandu Horsfield
- = Miletus; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 426; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 75
Gerydus Boisduval, [1836]; Hist. nat. Ins., Spec. gén. Lépid. 1: pl. 23 [= pl. 7C], f. 2; TS: Papilio symethus Cramer
- = Miletus; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 426; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 75
Archaeogerydus Fruhstorfer, 1915; in Seitz, Gross-Schmett Erde 9: 816; TS: Gerydus croton Doherty
- = Miletus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 75

Archaeogerydus; [CoL] ; [NHM-BMW]
Archaeogerydus (Section Miletus); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 426
Gerydus; [NHM-BMW]
Miletus; Eliot, 1961, Bull. Raffles Mus. 26: 154-177; [CoL] ; [iNaturalist] ; [NHM-BMW]
Miletus (Section Miletus); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 426
Miletus (Miletini); Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 75; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 116
Simoetheus; Boisduval, 1832, in d'Urville, Voy. Astrolabe (Faune ent. Pacif.) 1: 72 (missp.)
Symetha; [NHM-BMW]
Symethis; Guérin-Méneville, [1831], Voyage Coquille, 2 (2): pl. 18 (missp.); [CoL] ; [iNaturalist] ; [NHM-BMW]
Symethus; Guérin-Méneville, [1838], Voyage Coquille, 2 (2): 318 (missp.); [NHM-BMW]

1324x1220 (~162Kb) Thailand, Chon Buri Province, Pattaya environs, Ko Khram island, a bank of a pond in a spiny bamboo forest at a beach. 2nd February 2005, Photo © Oleg Kosterin

The exact identification of this species is still unknown, but tentatively assumed to belong into this group.

Larva on (aphids and coccids) [BMP]

The chinensis species-group

Chinensis; Eliot, 1961, Bull. Raffles Mus. 26: 154

Miletus chinensis Felder, 1862 -. See [maps] Common Brownie
- Miletus chinensis Felder, 1862; Verh. zool.-bot. Ges. Wien 12 (1/2): 488, no. 146; TL: Hong Kong
- Miletus chinensis; [MRS], 614; Eliot, 1961, Bull. Raffles Mus. 26: 154, 161; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; Huang & Xue, 2004, Neue Ent. Nachr. 57: 156 (note), f. 6, 7e, pl. 12a, f. 5; [CoL] ; [iNaturalist]
- M. c. chinensis Yunnan - SE.China, Hainan
  - Gerydus boisduvali chinensis; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Miletus chinensis chinensis; Eliot, 1961, Bull. Raffles Mus. 26: 162; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist]
- M. c. learchus C. & R. Felder, [1865] Indo-China, Siam, Tenasserim, N.Malaya
  - Miletus learchus C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 285, pl. 35, f. 37; TL: "Luzon" [error, = Cochin China]
  - Gerydus irroratus Druce, 1874; Proc. zool. Soc. Lond. 1874 (1): 106; TL: near Bangkok
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 162; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4
  - Miletus archilochus kelantanus Corbet, 1938; J. fed. Malay St. Mus. 18 (2): 248; TL: Kelantan, Malaya
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 162; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4
  - Gerydus boisduvali irroratus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Gerydus learchus learchus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 342
  - Miletus chinensis learchus; Eliot, 1961, Bull. Raffles Mus. 26: 162; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [BMP]: 220, pl. 33, f. 3-4; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
  1093x1177 (~438Kb) ♀ underside Cambodia, Koh Kong Province, Tatai Commune, Phum Doung village, 22nd March 2015, Photo © Oleg Kosterin
- M. c. assamensis (Doherty, 1891) Kumaon - Assam, Sikkim
  - Gerydys irroratus var. assamensis Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (1): [?] 37. pl. 1, f. 7; TL: Dhansiri Valley, Assam
  - Gerydys boisduvali milvius Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 309; TL: Sikkim
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 161; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4
  - Gerydys boisduvali assamensis; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Miletus chinensis assamensis; Eliot, 1961, Bull. Raffles Mus. 26: 161; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [Torben B. Larsen]; [CoL] ; [iNaturalist]
- M. c. longeana (de Nicéville, 1898) Manipur, Burma Long's Brownie
  - Gerydus longeana de Nicéville, 1898; J. Bombay nat. Hist. Soc. 12 (1): 141, pl. Z, f. 15-16; TL: Hsipau, North Shan States
  - Gerydus longeana; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 341
  - Miletus chinensis longeana; Eliot, 1961, Bull. Raffles Mus. 26: 162; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
Miletus croton (Doherty, 1889) -. See [maps]
- Gerydus croton Doherty, 1889; J. Asiat. Soc. Bengal 58 Pt.II (4): 439, pl. 23, f. 9; TL: Lower Tenasserim; India
- Gerydus croton; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 310
- Miletus croton; Eliot, 1961, Bull. Raffles Mus. 26: 154, 162; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist] ; [LEPIFORUM]
- M. c. croton Burma
  - Gerydus croton tavoyana Evans, 1932; Indian Butterflies (edn. 2): 210; TL: Tavoy, Burma
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 163; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4
  - Gerydus croton croton; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 310
  - Miletus croton croton; Eliot, 1961, Bull. Raffles Mus. 26: 163; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist] ; [Yutaka Inayoshi]
- M. c. corus Eliot, 1961 S.Shan States, NW.Siam
  - Miletus croton corus Eliot, 1961; Bull. Raffles Mus. 26: 162; TL: South Shan States, Kengtung District, Loi Mwe
  - Miletus croton corus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist] ; [Yutaka Inayoshi]
Miletus karennia (Evans, 1932) Karen Hills
- Gerydys croton karennia Evans, 1932; Indian Butterflies (edn. 2): 210; TL: Karen Hills
- Miletus croton karennia; Eliot, 1961, Bull. Raffles Mus. 26: 162
- Miletus croton karennius; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4
- Miletus karennia; [Torben B. Larsen, (note)]; [iNaturalist]
Miletus mallus (Fruhstorfer, 1913) -. See [maps]
- Gerydus croton mallus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 310; TL: S.Annam
- Miletus mallus; [MRS], 614; Eliot, 1961, Bull. Raffles Mus. 26: 154; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4, 78; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
- M. m. mallus Burma, Thailand, Laos, S.Vietnam
  - Miletus mallus mallus; Eliot, 1961, Bull. Raffles Mus. 26: 163; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist]
- M. m. shania (Evans, 1932) Burma
  - Gerydus gethusus shania Evans, 1932; Indian Butterflies (edn. 2): 211; TL: North Shan States
  - Miletus mallus shania; Eliot, 1961, Bull. Raffles Mus. 26: 163; [CoL] ; [iNaturalist]
  - Miletus mallus shanius; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [MRS], 614
- M. m. gethusus (Fruhstorfer, 1915) S.Yunnan, N.Vietnam, ...
  - Gerydus gethusus Fruhstorfer, 1915; Gross-Schmett. Erde 9: 816; TL: Chiem Hoa, Tonkins
    
    = Miletus mallus mallus; Eliot, 1961, Bull. Raffles Mus. 26: 163
  - Miletus mallus gethusus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; Huang & Xue, 2004, Neue Ent. Nachr. 57: 156 (note), pl. 12a, f. 3-4; [iNaturalist]
Miletus bannanus Huang & Xue, 2004 S.Yunnan. See [maps]
- Miletus bannanus Huang & Xue, 2004; Neue Ent. Nachr. 57: 155, pl. 12a, f. 1-2; TL: 20km W of Meng-la, Xi-shuang-ban-na, S.Yunnan, China
- Miletus bannanus; [iNaturalist]
Miletus gaesa (de Nicéville, 1895) -. See [maps]
- Gerydus gaesa de Nicéville, 1895; J. Bombay nat. Hist. Soc. 10 (1): 26, pl. S, f. 16; TL: NE.Sumatra
- Miletus gaesa; Eliot, 1961, Bull. Raffles Mus. 26: 154, 163; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
- M. g. gaesa Sumatra, Malaya
  - Gerydus boisduvali gaesa; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Miletus gaesa gaesa; Eliot, 1961, Bull. Raffles Mus. 26: 163; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [BMP]: 220, pl. 33, f. 1-2; [iNaturalist] ; [Yutaka Inayoshi]
- M. g. carrinas (Fruhstorfer, 1915) Borneo
  - Gerydus learchus carrinas Fruhstorfer, 1915; Gross-Schmett. Erde 9: 816; TL: S.Borneo
  - Miletus gaesa carrinas; Eliot, 1961, Bull. Raffles Mus. 26: 163; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
Miletus nymphis (Fruhstorfer, 1913) -. See [maps]
- Miletus nymphis; Eliot, 1961, Bull. Raffles Mus. 26: 154, 164; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [BOW]: pl. 178, f. 30 (text); [MRS], 614; [CoL] ; [iNaturalist]
- M. n. nymphis Fruhstorfer, 1913 W.Sumatra
  - Gerydus biggsi nymphis Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307, (♂ only) ; TL: W.Sumatra
  - Miletus nymphis nymphis; Eliot, 1961, Bull. Raffles Mus. 26: 164; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist]
- M. n. fictus Corbet, 1939 SE.Siam, Malaya, NE.Sumatra
  - Miletus fictus Corbet, 1939; Proc. R. ent. Soc. Lond. (B) 8 (2): 26; TL: Malaya
  - Miletus nymphis fictus; Eliot, 1961, Bull. Raffles Mus. 26: 164; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [BMP]: 219, pl. 33, f. 5-6; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
- M. n. porus Eliot, 1961 Burma
  - Miletus nymphis porus Eliot, 1961; Bull. Raffles Mus. 26: 164; TL: Karen Hills
  - Miletus nymphis porus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
- M. n. eneus Eliot, 1961 SE.Sumatra
  - Miletus nymphis eneus Eliot, 1961; Bull. Raffles Mus. 26: 164; TL: SE.Sumatra, Liwa, 900-1,400m
  - Miletus nymphis eneus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]

The zinckenii species-group

Zinckenii; Eliot, 1961, Bull. Raffles Mus. 26: 154

Miletus zinckenii C. & R. Felder, [1865] -. See [maps]
- Miletus zinckenii C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 284
- Gerydus zinckeni; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 247
- Miletus zinckenii; Eliot, 1961, Bull. Raffles Mus. 26: 154, 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [BOR], 546; [CoL] ; [iNaturalist]
- M. z. zinckenii Java
  - Gerydus zinckeni zinckeni; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 307
  - Miletus zinckenii zinckenii; Eliot, 1961, Bull. Raffles Mus. 26: 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist]
- M. z. improbus (Druce, 1896) Borneo
  - Gerydus improbus Druce, 1896; Proc. zool. Soc. Lond. 1896 (3): 651, pl. 29, f. 1-2; TL: N. Borneo, Kina Balu
  - Gerydus zinckeni improbus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 307
  - Miletus zinckenii improbus; Eliot, 1961, Bull. Raffles Mus. 26: 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; Takanami, 1989, Tyô to Ga 40 (1): 24; [CoL] ; [iNaturalist]
Miletus gopara (de Nicéville, 1890) -. See [maps]
- Gerydus gopara de Nicéville, 1890; Butts India Burmah Ceylon 3: 25; TL: Perak
  - = Gerydys biggsii; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 560
- Miletus gopara gopara; [BMP]: 221, pl. 33, f. 7-8
- Miletus gopara; Eliot, 1961, Bull. Raffles Mus. 26: 154, 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
- M. g. gopara Thailand, Malaysian Peninsula, Tioman, Singapore, Sumatra, Java
  - Gerydys biggsi f. denticulata Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307; TL: NE.Sumatra
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4
  - Miletus gopara gopara; Eliot, 1961, Bull. Raffles Mus. 26: 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [iNaturalist] ; [Yutaka Inayoshi]
- M. g. eustatius (Fruhstorfer, 1913) Borneo
  - Gerydus biggsi eustatius Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 308; TL: N. Borneo
  - Miletus gopara eustatius; Eliot, 1961, Bull. Raffles Mus. 26: 165; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
- M. g. artaxatus (Fruhstorfer, 1913) Java
  - Gerydus biggsi artaxatus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 308; TL: W.Java
  - Gerydus biggsi artaxatus f. oichalia Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 308; TL: W.Java
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Miletus gopara artaxatus; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. g. pardus Eliot, 1961 Sumatra
  - Miletus gopara pardus Eliot, 1961; Bull. Raffles Mus. 26: 165; TL: Sumatra, Lebong Tandai, 3,500'
  - Miletus gopara pardus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
Miletus valeus (Fruhstorfer, 1913) Malaya, NE.Sumatra. See [maps]
- Gerydus zinckeni valeus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307; TL: NE.Sumatra
- Gerydus zinckeni pallaxopas Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307; TL: Selangor
  - = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
- Miletus valeus; Eliot, 1961, Bull. Raffles Mus. 26: 154, 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BMP]: 221, pl. 33, f. 9; [CoL] ; [iNaturalist]
Miletus gaetulus (de Nicéville, 1894) -. See [maps]
- Gerydus gaetulus de Nicéville, 1894; J. Asiat. Soc. Bengal 63 Pt.II (1): 24, pl. 5, f. 12; TL: NE.Sumatra, Battak Mts.
- Gerydus gaetulus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 341
- Miletus gaetulus; Eliot, 1961, Bull. Raffles Mus. 26: 154, 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. g. gaetulus NE.Sumatra
  - Gerydus gaetulus gaetulus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 341
  - Miletus gaetulus gaetulus; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [iNaturalist]
- M. g. innocens (Druce, 1895) Borneo, S.Sumatra
  - Gerydus innocens Druce, 1895; Proc. zool. Soc. Lond. 1895 (3): 560, pl. 31, f. 4; TL: N.Borneo, Mt. Kina Balu
  - Gerydus gaetulus innocens; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 342
  - Miletus gaetulus innocens; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Takanami, 1992, Tyô to Ga 43 (1): 41 (note); [CoL] ; [iNaturalist]
- M. g. aphytis (Fruhstorfer, 1913) Nias
  - Gerydus gaetulus aphytis Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (11): 342; TL: Nias
  - Miletus gaetulus aphytis; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]

The boisduvali species-group

Boisduvali; Eliot, 1961, Bull. Raffles Mus. 26: 155

Miletus boisduvali Moore, [1858] -. See [maps] Common Brownie Boisduval's Brownie
- Miletus boisduvali Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 19, pl. 1a, f. 1; TL: Java
- Miletus boisduvali; Eliot, 1961, Bull. Raffles Mus. 26: 155, 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BOW]: pl. 178, f. 31; [MRS], 614; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
Larva preys on Ceratophis , Pseudococcus , (attended by Dolichoderus) Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117
- M. b. boisduvali Sundaland, Java, Sumatra, Borneo, Moluccas, New Guinea
  - Gerydus stygianus Butler, 1884; Ann. Mag. Nat. Hist. (5) 13 (75): 194; TL: Ternate
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6
  - Miletus chinensis var. ceramensis Ribbe, 1889; Dt. ent. Z. Iris 2 (2): 248, 202, pl. 5, f. 2; TL: Ceram
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Takanami, 1989, Tyô to Ga 40 (1): 47; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6
  - Gerydus boisduvali var. acragas Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2): 186; TL: Sumba
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus vincula Druce, 1895; Proc. zool. Soc. Lond. 1895 (3): 561, pl. 31, f. 9-10; TL: Borneo
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6
  - Gerydus buruensis Holland, 1900; Novit. Zool. 7 (1): 67; TL: Buru
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6
  - Gerydus boisduvali lombokianus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 309; TL: Lombok
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6
  - Gerydus boisduvali dossemus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 310; TL: Obi
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7
  - Gerydus boisduvali adeus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 310; TL: New Guinea, Fak Fak
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7
  - Gerydus boisduvali heraeon Fruhstorfer, 1915; Gross-Schmett. Erde 9: 818; TL: W.Borneo
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7
  - Gerydus courvoisieri Fruhstorfer, 1915; Z. wiss. Insektenbiol. 11 (9/10): 268; TL: W.Java, nr Buitenzorg
  - Gerydus courvoisieri Fruhstorfer, 1915; Gross-Schmett. Erde 9: 818; TL: Java
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7; Cassidy, 2013, Nachr. ent. Ver. Apollo, N.F. 34 (3): 141
  - Gerydus boisduvali oxylus Fruhstorfer, 1915; Gross-Schmett. Erde 9: 818; TL: Bawean
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7
  - Gerydus boisduvali acragas; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Gerydus boisduvali boisduvali; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Gerydus boisduvali buruensis; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 310
  - Gerydus boisduvali ceramensis; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Gerydus boisduvali stygianus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 310
  - Gerydus boisduvali vincula; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 309
  - Miletus boisduvali boisduvali; Eliot, 1961, Bull. Raffles Mus. 26: 166; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BMP]; [BAUR], 384; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 6; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [iNaturalist]
- M. b. diotrophes (Fruhstorfer, 1913) E.Celebes
  - Gerydus boisduvali diotrophes Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 309; TL: East Celebes
  - Miletus boisduvali diotrophes; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
- M. b. avitus (Fruhstorfer, 1915) Kai I.
  - Gerydus boisduvali avitus Fruhstorfer, 1915; Gross-Schmett. Erde 9: 818; TL: Key Is.
  - Miletus boisduvali avitus; Eliot, 1961, Bull. Raffles Mus. 26: 167; [BAUR], 384 (text); [CoL] ; [iNaturalist]
  - Milteus boisduvali avitus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
Miletus drucei (Semper, 1889) Philippines. See [maps]
- Gerydus drucei Semper, 1889; Reisen Philipp. (4): 162, pl. 31, f. 8-9; TL: Bohol
- Miletus drucei; Eliot, 1961, Bull. Raffles Mus. 26: 155, 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. d. drucei
  - Miletus philippus Staudinger, 1889; Dt. ent. Z. Iris 2 (1): 92, pl. 1, f. 2; TL: Palawan
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Takanami, 1989, Tyô to Ga 40 (1): 24
  - Gerydus boisduvali jacchus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 309; TL: Luzon
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus boisduvali paianius Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 309; TL: Mindoro
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus boisduvali epidurus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 309; TL: Palawan
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus biggsi drucei; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 308
  - Gerydus learchus philippus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 342
  - Gerydus philippus; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 561
  - Miletus drucei drucei; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [iNaturalist]
- M. d. metrovius (Fruhstorfer, 1913) Borneo
  - Gerydus biggsi metrovius Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 308; TL: N.Borneo
  - Gerydus courvoisieri phradimon Fruhstorfer, 1915; Z. wiss. Insektenbiol. 11 (9/10): 269; TL: N.Borneo
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Miletus drucei metrovius; Eliot, 1961, Bull. Raffles Mus. 26: 167; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
Miletus biggsii (Distant, 1884) Burma, Malaya. See [maps] Bigg's Brownie
- Gerydus biggsii Distant, 1884; Rhopalocera Malayana: 206, pl. 22, f. 12; TL: Malay Peninsula
- Gerydus biggsi; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 307
- Gerydus biggsii; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 561
- Miletus biggsi; [BOW]: pl. 178, f. 30
- Miletus biggsii biggsii; [BMP]: 220, pl. 33, f. 10-12
- Miletus biggsii; Eliot, 1961, Bull. Raffles Mus. 26: 155, 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. b. biggsii S.Burma, Thailand, Langkawi, Malay Peninsula, Tioman, Singapore, Borneo, Sumatra, Belitung, Weh
  - Gerydus biggsi f. atomaria Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307; TL: Rhio Archipelago
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus boisduvali hyllus Fruhstorfer, 1915; Gross-Schmett. Erde 9: 818; TL: NE.Sumatra
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus boisduvali xeragis Fruhstorfer, 1915; Gross-Schmett. Erde 9: 818; TL: Singapor
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus biggsi sebethus Fruhstorfer, 1915; Gross-Schmett. Erde 9: 819; TL: W.Borneo
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus biggsi extraneus Toxopeus, 1929; Tijdschr. Ent. 72 (3/4): 211; TL: Pulo Weh
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus biggsi biggsi; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 307
  - Miletus biggsii atomaria; [CoL]
  - Miletus biggsii biggsii; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [iNaturalist] ; [Yutaka Inayoshi]
  - Miletus biggsii extraneus; [CoL]
- M. b. niasicus (Fruhstorfer, 1913) Nias I., Batu I.
  - Gerydus biggsi niasicus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307; TL: Nias
  - Gerydus biggsi batunensis Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 307; TL: Batu Is.
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168
  - Miletus biggsii niasicus; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. b. albotignula (van Eecke, 1914) Simalur
  - Gerydus boisduvali albotignula van Eecke, 1914; Notes Leyden Mus. 36: 245; TL: Simalur
  - Gerydus boisduvali simalurensis Toxopeus, 1928; Tijdschr. Ent. 71: xxii; TL: Simalur
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Miletus biggsii albotignula; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. b. natunensis (Fruhstorfer, 1915) Natuna Is.
  - Gerydus biggsi natunensis Fruhstorfer, 1915; Gross-Schmett. Erde 9: 819 (Eecke i. l.); TL: Natuna Is.
  - Miletus biggsii natunensis; Eliot, 1961, Bull. Raffles Mus. 26: 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
  - Miletus natunensis; [CoL]
Miletus cellarius (Fruhstorfer, 1913) N.Borneo. See [maps]
- Gerydus biggsi cellarius Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (10): 308; TL: N.Borneo, Mt. Kina Balu
- Miletus cellarius; Eliot, 1961, Bull. Raffles Mus. 26: 155, 168; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]

The symethus species-group

Symethus; Eliot, 1961, Bull. Raffles Mus. 26: 155

Miletus symethus (Cramer, [1777]) -. See [maps] Great Brownie
- Papilio symethus Cramer, [1777]; Uitl. Kapellen 2 (9-16): 84, pl. 149, f. B, C; TL: "West Indies" [error, = Java]
- Gerydus symethus; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 559; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 243
- Miletus symethus; Eliot, 1961, Bull. Raffles Mus. 26: 155; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 78; [EBW]; [BOW]: pl. 178, f. 34; [CoL] ; [iNaturalist] ; [LEPIFORUM]
- M. s. symethus Java
  - Symetha pandu Horsfield, [1829]; Descr. Cat. lep. Ins. Mus. East India Coy (1): pl. 2, f. 2, 2a; TL: W.Java
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus symethus symethus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 243
  - Miletus symethus symethus; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [iNaturalist]
- M. s. petronius (Distant & Pryer, 1887) Burma, Thailand, Langkawi, Malay Peninsula, Singapore, Borneo, Bangka, Natuna, Belitung, Karimata
  - Gerydus petronius Distant & Pryer, 1887; Ann. Mag. nat. Hist. (5) 19 (112): 266; TL: N. Borneo, Sandakan
  - Gerydus symethus diopeithes Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 244; TL: Rhio Archipelago
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus symethus bangkanus Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 59; TL: Bangka
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus symethus hieropous Fruhstorfer, 1915; Gross-Schmett. Erde 9: 822; TL: Brunei
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus petronius; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 559
  - Gerydus symethus diopeithes; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 59
  - Gerydus symethus petronius; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 244; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 59
  - Miletus symethus bangkanus; [CoL]
  - Miletus symethus petronius; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BMP]: 221, pl. 33, f. 16-17; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
- M. s. perlucidus (Fruhstorfer, 1913) Bali, Lombok, E.Java
  - Gerydus symethus perlucidus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 243; TL: E.Java
  - Gerydus symethus megaris Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 244; TL: Lombok
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Miletus symethus megaris; [BOR], 546
  - Miletus symethus perlucidus; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. acampsis (Fruhstorfer, 1913) N.Sumatra
  - Gerydus symethus acampsis Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 243; TL: NE.Sumatra
  - Miletus symethus acampsis; Eliot, 1961, Bull. Raffles Mus. 26: 169; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. vespasianus (Fruhstorfer, 1913) Nias
  - Gerydus symethus vespasianus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 244; TL: Nias
  - Miletus symethus vespasianus; Eliot, 1961, Bull. Raffles Mus. 26: 170; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. edonus (Fruhstorfer, 1913) Palawan
  - Gerydus symethus edonus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 244; TL: Palawan
  - Gerydus symethus edonus; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 59
  - Miletus symethus edonus; Eliot, 1961, Bull. Raffles Mus. 26: 170; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. batuensis (Fruhstorfer, 1914) Batu Is.
  - Gerydus symethus batuensis Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 59; TL: Batu Is.
  - Miletus symethus batuensis; Eliot, 1961, Bull. Raffles Mus. 26: 170; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. philopator (Fruhstorfer, 1914) Philippines
  - Gerydus symethus philopator Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 59; TL: Mindoro
  - Miletus symethus philopator; Eliot, 1961, Bull. Raffles Mus. 26: 170; Kawazoe, 1973, Tyô to Ga 24 (4): 93; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. hierophantes (Fruhstorfer, 1915) N.Borneo, Sulu Is.
  - Gerydus symethus hierophantes Fruhstorfer, 1915; Gross-Schmett. Erde 9: 822; TL: Sulu Is.
  - Miletus symethus hierophantes; Eliot, 1961, Bull. Raffles Mus. 26: 170; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. nuctus Eliot, 1961 Sumatra
  - Miletus symethus nuctus Eliot, 1961; Bull. Raffles Mus. 26: 169; TL: SE.Sumatra, Liwa, 900-1400m
  - Miletus symethus nuctus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. s. solitarius Okubo, 1983 Tioman
  - Miletus ancon solitaria Okubo, 1983; Tyô to Ga 33 (3,4): 176, f. 27-28; TL: Lalang-Tekek, Tioman
  - Miletus symethus solitarius; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 78; [iNaturalist]
- M. s. phantus Eliot, 1986 Philippines (Marinduque, Luzon, ...)
  - Miletus symethus phantus Eliot, 1986; Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 78; TL: Philippines, Marinduque
  - Miletus symethus phantus; [CoL] ; [iNaturalist]
Miletus gallus (de Nicéville, 1894) Thailand, Borneo. See [maps]
- Gerydus gallus de Nicéville, 1894; J. Asiat. Soc. Bengal 63 Pt.II (1): 25, pl. 5, f. 11; TL: NE.Sumatra, Battak Mts
- Gerydus gallus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 341
- Miletus gallus; Eliot, 1961, Bull. Raffles Mus. 26: 155, 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. g. gallus Malaya, N.Sumatra
  - Miletus gallus gallus; Eliot, 1961, Bull. Raffles Mus. 26: 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BMP]: 220, pl. 33, f. 14-15; [iNaturalist] ; [Yutaka Inayoshi]
- M. g. leucocyon (Toxopeus, 1940) Java
  - Gerydon gallus leucocyon Toxopeus, 1940; Ent. Meded. Ned.-Ind. 6: 15; TL: Java
  - Miletus gallus leucocyon; Eliot, 1961, Bull. Raffles Mus. 26: 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
Miletus heracleion (Doherty, 1891) -. See [maps]
- Gerydus heracleion Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (1): 36; TL: Perak
- Gerydus heracleion; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 310
- Miletus heracleion; Eliot, 1961, Bull. Raffles Mus. 26: 155, 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4, 80; [CoL] ; [iNaturalist]
- M. h. heracleion Malaya
  - Miletus heracleion heracleion; Eliot, 1961, Bull. Raffles Mus. 26: 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [BMP]: 220, pl. 61, f. 10 (m holotype), pl. 63, f. 5 (m holotype un.); [iNaturalist]
- M. h. arion Eliot, 1961 Borneooo, Pulo Laut
  - Miletus heracleion arion Eliot, 1961; Bull. Raffles Mus. 26: 171; TL: Sabah
  - Miletus heracleion arion; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 4; [CoL] ; [iNaturalist]
Miletus ancon (Doherty, 1889) -. See [maps]
- Gerydus ancon Doherty, 1889; J. Asiat. Soc. Bengal 58 Pt.II (4): 438, pl. 23, f. 8; TL: Tavoy
- Gerydus ancon; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 246
- Miletus ancon; Eliot, 1961, Bull. Raffles Mus. 26: 155, 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BOW]: pl. 178, f. 28; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
- M. a. ancon Burma (Shan States, Karen Hills, Tavoy)
  - Gerydus ancon ancon; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 246
  - Miletus ancon ancon; Eliot, 1961, Bull. Raffles Mus. 26: 171; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BOR], 546; [iNaturalist] ; [Yutaka Inayoshi]
- M. a. gigas (Druce, 1895) Borneo
  - Gerydus gigas Druce, 1895; Proc. zool. Soc. Lond. 1895 (3): 559, pl. 31, f. 3; TL: N.Borneo, Mt. Kina Balu
  - Gerydus ancon anconides Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 246; TL: Sarawak
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus gigas; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 341
  - Miletus ancon gigas; Eliot, 1961, Bull. Raffles Mus. 26: 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Takanami, 1992, Tyô to Ga 43 (1): 41 (note); [CoL] ; [iNaturalist]
Miletus archilochus (Fruhstorfer, 1913) Vietnam, ...?. See [maps]
- Gerydus archilochus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 247; TL: Central Tonkin
- Miletus archilochus; Eliot, 1961, Bull. Raffles Mus. 26: 155; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BOW]: pl. 178, f. 29; [MRS], 614; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
- M. a. archilochus
  - Miletus archilochus archilochus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [iNaturalist]
- M. a. siamensis (Godfrey, 1916) Thailand, Laos, Vietnam
  - Gerydus ancon siamensis Godfrey, 1916; J. nat. Hist. Soc. Siam 2 (2): 134; TL: Thailand
  - Elkalyce siamensis; [CoL]
  - Miletus ancon siamensis; Eliot, 1961, Bull. Raffles Mus. 26: 171; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
  - Miletus archilochus siamensis; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 79; [iNaturalist]
  807x806 (~224Kb) ♀ underside Cambodia, Siem Reap Province, Phnom Kulen Mts., a brook, the left Siem Reap River tributary, 1.2 km SE of Preah Ang Thom, 30th June 2018, Photo © Oleg Kosterin
- M. a. guangxiensis Li, 1994 Guangxi
  - Miletus archilochus guangxiensis Li, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 767, 614; TL: Longzhou 240m, Guanxi
  - Miletus archilochus guangxiensis; [iNaturalist]
Miletus gigantes (de Nicéville, 1894) Malaya, Sumatra. See [maps]
- Gerydus gigantes de Nicéville, 1894; J. Asiat. Soc. Bengal 63 Pt.II (1): 23, pl. 4, f. 1, 13; TL: NE.Sumatra, Battak Mts.
- Gerydus gigantes; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (11): 341
- Miletus ancon gigantes; Eliot, 1961, Bull. Raffles Mus. 26: 171; [CoL]
- Miletus gigantes; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 79; [BMP]: 221, pl. 33, f. 13; [BOR], 546; [BOW]: pl. 178, f. 28 (text); [iNaturalist] ; [Yutaka Inayoshi]
Miletus atimonicus (Murayama & Okamura, 1973) Philippines (Luzon). See [maps]
- Miletus sumethus[sic] atimonicus Murayama & Okamura, 1973; Tyô to Ga 24 (1): 23, f. 45-46; TL: Philippines, Luzon, Atimonan
  - = Miletus symethus philopator; Kawazoe, 1973, Tyô to Ga 24 (4): 93
- Miletus atimonicus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 79; [CoL] ; [iNaturalist]
Miletus celinus Eliot, 1961 S.Celebes. See [maps]
- Miletus celinus Eliot, 1961; Bull. Raffles Mus. 26: 155, 175; TL: S.Celebes
- Miletus celinus; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BOW]: pl. 178, f. 33 (text); Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 8; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
Miletus rosei Cassidy, 1995 N.Sulawesi. See [maps]
- Miletus rosei Cassidy, 1995; Trans. lepid. Soc. Japan 46 (1): 8, f. 18-21; TL: North Sulawesi, Dumoga Bone National Park, Toraut District, 1000m
- Miletus rosei; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [iNaturalist]
Miletus takanamii Eliot, 1986 Philippines (Mindanao). See [maps]
- Miletus takanamii Eliot, 1986; Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 83, f. 102-103; TL: Philippines, Mindanao, Tandag, Surigao
- Miletus takanamii; [CoL] ; [iNaturalist]
Miletus leos (Guérin-Méneville, 1829) Moluccas, New Guinea. See [maps]
- Simaethus leos Guérin-Méneville, 1829; Voyage Coquille, 2 (2): pl. 18, f. 8 ♀; TL: Buru
- Gerydus leos; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 244
- Miletus leos catoleucus; [CoL]
- Miletus leos; Eliot, 1961, Bull. Raffles Mus. 26: 155, 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BOW]: pl. 178, f. 32; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
- M. l. leos S.Moluccas(Serang, Ambon, Buru)
  - Gerydus boisduvali Butler, 1884; Ann. Mag. Nat. Hist. (5) 13 (75): 194 (preocc.); TL: Amboina
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos meronus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 245; TL: Ceram
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos amphiarus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 245; TL: "Banggai" [error?]
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos gardineri Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 59; TL: Amboina
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus boisduvali; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (10): 308
  - Miletus leos leos; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BAUR], 384; [iNaturalist]
- M. l. maximus (Holland, 1890) Celebes, Bouton, Sangir, Talaut, Buton, Muna, Banggai
  - Gerydus maximus Holland, 1890; ; TL: S.Celebes
  - Gerydus leos maximus f. divisa Fruhstorfer, 1913; Z. wiss. Insektenbiol.: (?) ; TL: S.Celebes
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 173; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos sarus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 245; TL: E.Celebes
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 173; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7
  - Gerydus symethus vaneeckei Toxopeus, 1930; De soorte .. Plaats. en Tijd.: 112; TL: NE.Celebes, Minahassa
    
    = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7
  - Gerydus leos maximus; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 245
  - Miletus leos maximus; Eliot, 1961, Bull. Raffles Mus. 26: 173; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 7; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
  - Miletus leos vaneeckei; Eliot, 1961, Bull. Raffles Mus. 26: 173; [CoL]
  - Miletus leos vaneecki[sic]; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
- M. l. teos (Doherty, 1891) Sumba, ?Sumbawa
  - Gerydus teos Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2): 185; TL: Sumba
  - Gerydus leos teos; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 245
  - Miletus leos teos; Eliot, 1961, Bull. Raffles Mus. 26: 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. l. florensis (Fruhstorfer, 1913) Flores, Pura, Adonara, Alor, ?Sumbawa
  - Gerydus leos florensis Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 244; TL: Flores
  - Gerydus leos eulus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 244; TL: Sumbawa
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos eulus; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 59
  - Miletus leos florensis; Eliot, 1961, Bull. Raffles Mus. 26: 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [CoL] ; [iNaturalist]
- M. l. catoleucos (Fruhstorfer, 1913) Salayer
  - Gerydus leos catoleucos Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 246; TL: Salayer
  - Miletus leos catoleucos; Eliot, 1961, Bull. Raffles Mus. 26: 173; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 8; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [iNaturalist]
- M. l. mangolicus (Fruhstorfer, 1913) Sula Is.
  - Gerydus leos mangolicus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 245; TL: Sula Mangoli
  - Miletus leos mangolicus; Eliot, 1961, Bull. Raffles Mus. 26: 173; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
- M. l. virtus (Fruhstorfer, 1913) N.Moluccas (Bachan, Ternate, Halmahera)
  - Gerydus leos virtus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 245; TL: Batchian
  - Gerydus leos pentheus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 245 [wet season form]; TL: Halmaheira
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174 (form); Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos virtus f. virtus; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 60
  - Miletus leos virtus; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BAUR], 385 (text); [CoL] ; [iNaturalist]
  - Miletus leos virtus f. pentheus; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 60
- M. l. tellus (Fruhstorfer, 1913) Wetar?, Timor?
  - Gerydus ancon tellus Fruhstorfer, 1913; Z. wiss. Insektenbiol. 9 (8/9): 246; TL: "E.Java" [?, = Wetter?]
  - Gerydus leos tellus; Fruhstorfer, 1914, Z. wiss. Insektenbiol. 10 (2): 59
  - Miletus leos tellus; Eliot, 1961, Bull. Raffles Mus. 26: 172; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BAUR], 385 (text); Cassidy, 1995, Trans. lepid. Soc. Japan 46 (1): 8; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 117; [CoL] ; [iNaturalist]
- M. l. aronicus (Fruhstorfer, 1914) Aru, Misool, Waigeu, New Guinea
  - Gerydus leos aronicus Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 60; TL: Aru Is.
  - Gerydus leos rex Fruhstorfer, 1913; Z. wiss. Insektenbiol.: ?; TL: Waigeu
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174
  - Gerydus leos nineyanus Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 60; TL: New Guinea, Niney
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos acrisius Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 60; TL: New Guinea, Kapaur
    
    = ; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5
  - Gerydus leos rex; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 245
  - Miletus leos aronicus; Eliot, 1961, Bull. Raffles Mus. 26: 174; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5; [BAUR], 385 (text); [CoL] ; [iNaturalist]

The melanion species-group

Melanion; Eliot, 1961, Bull. Raffles Mus. 26: 156

Miletus melanion C. & R. Felder, [1865] -. See [maps]
- Miletus melanion C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 284, pl. 35, f. 32-33
- Miletus melanion ♂-f. albiguttatus Eliot, 1986; Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 80, f. 98; TL: Philippines, Luzon
- Gerydus melanion; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 246
- Miletus melanion bazilanus; [CoL]
- Miletus melanion vitelianus; [CoL]
- Miletus melanion; Eliot, 1961, Bull. Raffles Mus. 26: 156, 175; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 80; [BOW]: pl. 178, f. 33; [CoL] ; [iNaturalist]
- M. m. melanion Philippines
  - Gerydus melanion melanion; Fruhstorfer, 1913, Z. wiss. Insektenbiol. 9 (8/9): 246
  - Miletus melanion melanion; Eliot, 1961, Bull. Raffles Mus. 26: 175; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 81; [iNaturalist]
- M. m. euphranor (Fruhstorfer, 1914) Philippines (Mindoro)
  - Gerydus melanion euphranor Fruhstorfer, 1914; Z. wiss. Insektenbiol. 10 (2): 60; TL: Mindoro
    
    = Miletus melanion melanion; Eliot, 1961, Bull. Raffles Mus. 26: 175
  - Miletus melanion euphranor; Eliot, 1986, Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1): 5, 82; [iNaturalist]

Unmatched external taxa

Miletus ageles; [CoL]
Miletus agragas; [CoL]
Miletus architas; [CoL]
Miletus cataleucus; [CoL]
Miletus cratexas; [CoL]
Miletus docus; [CoL]
Miletus dryone; [CoL]
Miletus floresianus; [CoL]
Miletus milesius; [CoL]
Miletus modestus; [CoL]
Miletus moorei; [CoL]
Miletus plautus; [CoL]
Miletus poltinus; [CoL]
Miletus polydetus; [CoL]
Miletus pyane; [CoL]
Miletus ribbei; [CoL]
Miletus shania; [Yutaka Inayoshi]
Miletus sinensis; [CoL]
Miletus stotharti; [CoL]
Miletus tayoyana; [CoL]

28.2.2022 (27)

References:

[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[RAFFLES]
Bulletin of the Raffles Museum (1928-1960); Bulletin of the National Museum (1970); The Raffles Bulletin of Zoology (1988-2013)
[Torben B. Larsen]
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Boisduval, 1832
Voyage de découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les années 1826-1827-1828-1829, sous le commandément de M. J. Dumont d'Urville. Faune entomologique de l'Océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Lépidoptères in d'Urville, Voy. Astrolabe (Faune ent. Pacif.) 1 : 1-267, pl. 1-5
Boisduval, [1836]
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, 1884
The Lepidoptera collected during the recent expedition of H.M.S. "Challenger", Part II Ann. Mag. Nat. Hist. (5) 13 (75) : 183-203
Cassidy, 1995
On the Miletini (Lepidoptera, Lycaenidae) of the Sulawesi Region Trans. lepid. Soc. Japan 46 (1) : 1-12
Cassidy, 2013
On some type specimens of Lycaenidae from South East Asia (Lepidoptera) Nachr. ent. Ver. Apollo, N.F. 34 (3) : 137-144
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
Corbet, 1938
Observations on certain Species of Rhopalocera from the Malay Peninsula J. fed. Malay St. Mus. 18 (2) : 242-262
Corbet, 1939
A revision of the Malayan species of Miletus Hübner, (= Gerydys Boisduval) (Lepidoptera: Lycaenidae) Proc. R. ent. Soc. Lond. (B) 8 (2) : 25-32
[BMP]; Corbet & Pendlebury, 1992
The butterflies of the Malay Peninsula. Fourth Edition revised by J. N. Eliot with plates by Bernard D'Abrera Butterflies Malay Peninsula (4th edn) : 1-595, pl. 1-69
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
[BAUR]; D'Abrera, 1977
Butterflies of the Australian Region (2nd edn) Butts. Aust. Region: 1-416
[BOR]; D'Abrera, 1982-1986
Butterflies of the Oriental Region I-III
de Nicéville, 1894
On new and little-known butterflies from the Indo-Malayan region J. Asiat. Soc. Bengal 63 Pt.II (1) : 1-59, pl. 1-5
de Nicéville, 1895
On new and little-known Butterflies from the Indo-Malayan Region J. Bombay nat. Hist. Soc. 10 (1): 13-40, (2): 169-194
de Nicéville, 1898
On New and Little-Known Butterflies from the Indo-Malayan, Austro-Malayan and Australian Regions J. Bombay nat. Hist. Soc. 12 (1) : 131-161, pl. X-Z,AA
Distant, 1884
Rhopalocera Malayana: a Description of the Butterflies of the Malay Peninsula Rhopalocera Malayana : 1-481, pl. 1-27,27A,27B,28-44
Distant & Pryer, 1887
On the Rhopalocera of Northern Borneo. Part I & II Ann. Mag. nat. Hist. (5) 19 (109): 41-56, (112): 264-275
Doherty, 1889
On certain Lycaenidae from lower Tenasserim J. Asiat. Soc. Bengal 58 Pt.II (4) : 409-440, pl. 23
Doherty, 1891
New and rare Indian Lycaenidae J. Asiat. Soc. Bengal 60 Pt.II (1) : 32-38, pl. 1
Doherty, 1891
The Butterflies of Sumba and Sumbawa, with some Account of the Island of Sumba J. Asiat. Soc. Bengal 60 Pt.II (2) : 141-197, pl. 2
Druce, 1874
A List of the Lepidopterous Insects collected by Mr. L. Layard at Chentaboon and Nahconchaisee, Siam, with Descriptions of new Species Proc. zool. Soc. Lond. 1874 (1) : 102-109, pl. 16
Druce, 1895
A Monograph of the Bornean Lycaenidae Proc. zool. Soc. Lond. 1895 (3) : 556-627, pl. 31-34
Druce, 1896
Further Contributions to our Knowledge of the Bornean Lycaenidae Proc. zool. Soc. Lond. 1896 (3) : 650-683, pl. 29-31
Eliot, 1961
An analysis of the genus Miletus Hübner (Lepidoptera: Lycaenidae) Bull. Raffles Mus. 26 : 154-177
Eliot, 1973
The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 373-505 (162fgs), pl. 1-6
Eliot, 1986
A review of the Miletini (Lepidoptera: Lycaenidae) Bull. Br. Mus. nat. Hist. (Ent. ) 53 (1) : 1-105, 108fgs
van Eecke, 1914
Studien über Indo-Australische Lepidopteren, Fauna Simalurensis Notes Leyden Mus. 36 : 193-258, pl. 4
Evans, 1932
The Identification of Indian Butterflies (edn. 2) Indian Butterflies (edn. 2) : iii-x,1-454, pl. 1-32
Felder, 1862
Verzeichniss der von den Naturforschern der k. k. Fregatte Novara gesammelten Macropepidoteren Verh. zool.-bot. Ges. Wien 12 (1/2) : 473-496
C. & R. Felder, [1865]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fruhstorfer, 1913; Fruhstorfer, 1914; Fruhstorfer, 1915
Übersicht der Gerydinae und Diagnosen neuer oder verkannter Formen (Lep., Lyc.) Z. wiss. Insektenbiol. 9 (8/9): 242-247 (1 September), 9 (10): 307-310 (15 October), 9 (11): 341-344 (15 November), 9 (12): 367-371 (15 December), 10 (1): 20-25 (20 January), 10 (2): 59-63 (15 February), 11 (9/10): 267-269, f. 1-4 (20 November)
Fruhstorfer, 1915
II. Subfamilie: Gerydinae in Seitz, Gross-Schmett Erde 9 : 803-808 (24 April, 1914), : 809-823 (4 August, 1915)
Godfrey, 1916
The Butterflies of Siam J. nat. Hist. Soc. Siam 2 (2) : 106-147
Guérin-Méneville, 1829; Guérin-Méneville, [1831]; Guérin-Méneville, [1838]
Crustacés, arachnides et insectes. 319pp, 22pls [lepidoptera 271-286] Voyage Coquille, 2 (2) : pl. 13-17 (1830), : pl. 14bis (1831), : pl. 18-19 (1831), : 271-286, 316-319 [index?] (1838)
Holland, 1900
The Lepidoptera of Buru. (1) Rhopalocera, (2) Heterocera Novit. Zool. 7 (1): 54-86, (3): 555-591
Horsfield, [1828]; Horsfield, [1829]
Descriptive Catalogue of the Lepidopterous Insects contained in the Museum of the Horourable East-India Company, illustrated by coloured figures of new species Descr. Cat. lep. Ins. Mus. East India Coy (1): 1-80, pl. 1-4 (1828), (2): 81-144, pl. 5-8, (expl.) [1-4] (1829)
Huang & Xue, 2004
A new species of Miletus from the extreme south of Yunnan, China Neue Ent. Nachr. 57: 155-160, pl. 12a
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Kawazoe, 1973
A revisional note on some Philippine Lycaenidae and Hesperiidae reported by Murayama and Okamura Tyô to Ga 24 (4): 91-98
[BOW]; Lewis, 1974
Butterflies of the World
de Nicéville in Marshall & de Nicéville, 1890
The Butterflies of India, Burmah and Ceylon, a descriptive Handbook of all the known Species of rhopalocerous Lepidoptera inhabiting that Region, with Notices of allied Species occurring in the neighbouring Countries along the Border, with numerous Illustrations. Vol. 3 Butts India Burmah Ceylon 3 : 1-503, pl. 25-29
Moore, [1858]
A Catalogue of the Lepidopterous Insects in the Museum of the Hon. East-India Company in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: : 1-278, pl. 1-12,1a,2a,3a,4a,5a,6a
Murayama & Okamura, 1973
Butterflies of Luzon Island in the Philippines, with descriptions of new species and subspecies Tyô to Ga 24 (1) : 10-25, 48 fgs
Okubo, 1983
Butterflies of Tioman Island, West Malaysia, with the description of new species Tyô to Ga 33 (3,4): 168-184
Ribbe, 1889
Beiträge zur Lepidopteren-Faun von Gross-Ceram Dt. ent. Z. Iris 2 (2) : 187-265, pl. 5
Fruhstorfer in Seitz, 1915
Die Großschmetterlinge des Indo-australischen Faunengebietes Gross-Schmett. Erde 9 : 1-766, : 767-1197
Semper, 1889
Die Schmetterlinge der Philippinischen Inseln. Reisen im Archipel der Philippinen Reisen Philipp. (1): 1-46, pl. 1-8, A (1886), (2): 47-86, pl. 9-16 (1887), (3): 87-134, pl. 17-24 (1888), (4): 135-174, pl. 25-32 (1889), (5): 175-238, pl. 33-38, B (1890), (6): 239-270, pl. 39-46 (1891), (7): 271-380, pl. 47-49 [iv] (1892), : 381-728
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Staudinger, 1889
Lepidoptera der Insel Palawan Dt. ent. Z. Iris 2 (1) : 3-180, pl. 1-2
Takanami, 1989
On some type specimens of Lycaenidae from South East Asia Tyô to Ga 40 (1) : 23-80
Takanami, 1992
An additional catalogue on type specimens of Lycaenidae (Lepidoptera) from South East Asia, preserved in Museum fur Naturkunde der Humbold-Universitat zu Berlin Tyô to Ga 43 (1) : 35-46
Toxopeus, 1929
Beschreibung einiger Schmetterlinge (Riodinidae und Lycaenidae) von Pulu Weh bei Sumatra. (Lycaenidae Australasiae V) Tijdschr. Ent. 72 (3/4) : 204-214
Toxopeus, 1930
De soort als functie van plaats en tijd getoetst aan de Lycaenidae van het Australaziatisch gebied De soorte .. Plaats. en Tijd.: 198pp, 4pls, 17fgs
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.

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