Hepialus

Hepialus Fabricius, 1775

Hepialus Fabricius, 1775; Syst. Ent.: 589; TS: Phalaena humuli Linnaeus
Hepiolus Illiger, 1801; Mag. Insektenk. (Illiger) 1: 138 (emend.)
- = Hepialus; [NACL], 2; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list)
Epialus [= Hepialus] Agassiz, 1847; Nom. zool. (Index univ.): 140, 178 (emend.)
- = Hepialus; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list)
Epiolus Agassiz, 1847; Nom. zool. (Index univ.): 141, 178 (emend.)
Epialus Lederer, 1852; (emend.)

Epialus; [NHM-BMW]
Epiolus; [NHM-BMW]
Hegialus; [NHM-BMW]
Hegialus[sic, Hepialus]; Fabricius, 1787, Mantissa Insectorum 2: 134
Hepialus; Godman & Salvin, 1887, Biol. centr.-amer., Lep. Heterocera 1: 231; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Hepialus (Hepialidae); [NACL], 2; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 138
Hepiolus; [NHM-BMW]
Trepialus; [NHM-BMW]
Trepialus[sic, Hepialus]; Latreille, 1805, Hist. nat. Crust. Ins. 14: 174

Hepialus humuli (Linnaeus, 1758)

EU. See [maps] Großer Hopfen-Wurzelbohrer Ghost Moth Geistermotte Hopfenspinner Humalaperho

Phalaena (Noctua) humuli Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 508; TL: Europe
humulator Haworth, 1802
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list)
Hepialus thulensis Newman, 1865; Entomologist 2: 162; TL: Shetlands
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 139
Hepialus humuli v. hethlandica Staudinger, 1871; Cat. Lepid. eur. Faunegeb. (Edn. 2): 60
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 139
ab. rosea Petersen, 1902; Beitr. Kunde Est.Liv- Kurlands 1902: 167; TL: Estonia
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 139
Hepiolus humuli ab. albida Spuler, 1910; Schmett. Eur. 2: 484
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepiolus humuli ab. thulea Spuler, 1910; Schmett. Eur. 2: 485
- = ; Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli f. azuga Pfitzner, 1912; Gross-Schmett. Erde 2: 434 (Caradja)
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli f. grandis Pfitzner, 1912; Gross-Schmett. Erde 2: 434
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli ab. dannenbergi Stephan, 1923; Soc. ent. 38: 46; TL: Europe
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli ab. pusillus Stephan, 1923; Soc. ent. 38: 46; TL: Europe
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli ab. ♂ rufomaculata Lempke, 1938; Tijdschr. Ent. 82: 303; TL: Holland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli ab. roseoornata Bytinski-Salz, 1939; Ent. Rec. 51: 81; TL: Tavistock, Great Britain
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli thulensis f. uniformis Bytinski-Salz, 1939; Ent. Rec. 51: 82; TL: Shetland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli thulensis f. albida Bytinski-Salz, 1939; Ent. Rec. 51: 82; TL: Shetland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
f. faeroensis Dahl, 1954; Fródsskaparrit 3: 134, 153; TL: Faroes
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
ab. fumosa Cockayne, 1955; Ent. Gazette 6: 5; TL: England, Shetland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
ab. radiata Cockayne, 1955; Ent. Gazette 6: 5; TL: England, Shetland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli f. ♀ postrufescens Lempke, 1961; Tijdschr. Ent. 104 (8): 180; TL: Holland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
Hepialus humuli f. ♀ postnigrescens Lempke, 1961; Tijdschr. Ent. 104 (8): 180; TL: Holland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140
f. griseomaculata van Wisselingh, 1965; Ent. Ber. 25: 146; TL: Holland
- = ; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 140

Hepialus humuli azuga; [LepIndex]
Hepialus humuli faeroensis; [LepIndex]
Hepialus humuli grandis; [LepIndex]
Hepialus humuli griseomaculata; [LepIndex]
Hepialus humuli postnigrescens; [LepIndex]
Hepialus humuli postrufescens; [LepIndex]
Hepialus humuli thulensis; [LepIndex]
Hepialus humuli; Nielsen, Robinson & Wagner, 2000, J. Nat. Hist. 34: 849 (list); Zou, Liu & Zhang, 2010, J. Hun. Univ. Sci. Tech. 25 (1): 117; Grehan, Mielke, Turner & Nielsen, 2023, ZooNova 28: 139; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [SVF]
Hepiolus[sic] humuli; [SPRK], 157

698x472 (~104Kb) larva Germany, Baden-Württemberg, Ötisheim, Scharrkessel (8°49'E, 48°58'N, 320m), 01.11.2000, Photo © Karl Hofsäß	510x657 (~34Kb) ♀ Germany, Baden-Württemberg, Schönenberg, Siedlungsbereich (8°49'E, 48°57'N, 230m), 02.06.2000, Photo © Karl Hofsäß	1024x448 (~90Kb) ♂ Russia, Moscow area, 17.06.2007, Photo © D. Smirnov
561x702 (~79Kb) ♂ Russia, Moscow area, 17.06.2007, Photo © D. Smirnov	902x482 (~84Kb) ♀ Russia, Moscow area, 12.06.2007, Photo © D. Smirnov	identification uncertain 857x337 (~55Kb) larva Russia, Moscow area, 30.5.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
700x502 (~44Kb) FINLAND: Ka: Virolahti, 671:53, m 10.7.1976, f 25-1.7.1983, Markku Savela leg.

References:

[BCA] electronic
BIOLOGIA CENTRALI-AMERICANA
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[Karl Hofsäß] Karl Hofsaess
e-mail: Karl Hofsäß
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Agassiz, 1847
Nomenclatoris zoologici. Index Universalis Nom. zool. (Index univ.) : 1-393
Bytinski-Salz, 1939
New and little known forms of Hepialus mostly from Great Britain Ent. Rec. 51 : 81-85
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus Syst. Ent. : 1-832
Fabricius, 1787
Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus Mantissa Insectorum 2 : 1-382
Godman & Salvin, 1887
Biologia Centrali-Americana; or Contributions to the Knowledge of the Fauna of Mexico and Central America. Zoology. Lepidoptera. Heterocera Biol. centr.-amer., Lep. Heterocera 1: 1-490, 3: pl. 1-101
Grehan, Mielke, Turner & Nielsen, 2023
A revised world catalogue of Ghost Moths (Lepidoptera: Hepialidae) with taxonomic and biological annotations ZooNova 28 : 1-337
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Lempke, 1938
Catalogus der Nederlandsche Macrolepidoptera. III Tijdschr. Ent. 82 : 242-304
Lempke, 1961
Catalogus der Nederlandse Macrolepitoptera (Achtse Supplement) Tijdschr. Ent. 104 (8) : 111-186, pl. 9-12
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Newman, 1865
Singular Geographical Race of Hepialus Humuli Entomologist 2 : 162
Nielsen, Robinson & Wagner, 2000
Ghostmoths of the world: a global inventory and bibliography of the Exoporia J. Nat. Hist. 34 : 823-878
Pfitzner in Seitz, 1912
Die Gross-Schmetterlinge des Palaearktischen Faunengebietes. 2. Die Palaearktischen Spinner & Schwärmer Gross-Schmett. Erde 2 : 1-479 [Macrolep.], pl. 1-56
[SPRK]; Seppänen, 1970
Suomen suurperhostoukkien ravintokasvit Animalia Fennica 14
Spuler, 1910
Die Schmetterlinge Europas. Dritte Auflage II. Band Schmett. Eur. 2: : 1-523
Staudinger in Staudinger & Wocke, 1871
Catalog der Lepidopteren des europaeischen Faunegebiets (Edn. 2): - Cat. Lepid. eur. Faunegeb. (Edn. 2) : 1-421, : 422-426 (Corrigenda et Addenda)
Stephan, 1923
Hepialus humuli L., die Geistermotte Soc. ent. 38 : 45-46
van Wisselingh, 1965
Macrolepidoptera in 1964 Ent. Ber. 25 : 145-146
Zou, Liu & Zhang, 2010
Revision of taxonomic system of the genus Hepialus (Lepidoptera, Hepialidae) currently adopted in China J. Hun. Univ. Sci. Tech. 25 (1) : 114-120

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