Catopsilia

Catopsilia Hübner, [1819] Emigrants

Catopsilia Hübner, [1819]; Verz. bek. Schmett. (7): 98; TS: Papilio crocale Cramer
Murtia Hübner, [1819]; Verz. bek. Schmett. (7): 98; TS: Mancipium minna Hübner
- = Catopsilia; [AfrL]

Catopsilia; Winhard, 2000, Butterflies of the world 10: 12; [CoL] ; [iNaturalist] ; [NHM-BMW]
Catopsilia (Coliadinae); [AfrL]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 100
Catopsilia (Coliadini); Zhang, Cong, Shen, Opler & Grishin, 2021, Taxonomic Rep. 9 (3): 7
Catopsilia (Pierinae); Moore, [1881], Lepid. Ceylon 1 (3): 121
Murtia; [NHM-BMW]

769x513 (~135Kb) Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko

Catopsilia pomona (Fabricius, 1775)

Ceylon, India, Burma, Malaya, New Guinea, Australia. See [maps] Lemon Migrant Lemon Emigrant Common Emigrant

Papilio pomona Fabricius, 1775; Syst. Ent.: 479, no. 158; TL: New Holland
Papilio crocale Cramer, [1775]; Uitl. Kapellen 1 (1-7): 87, pl. 55, f. C, D; TL: East Indies
Papilio jugurtha Cramer, [1777]; Uitl. Kapellen 2 (9-16): 138, pl. 187, f. E, F; TL: Coromandel
Papilio nigropunctatus Goeze, 1779; Ent. Beyträge 3 (1): 185
Papilio catilla Cramer, [1779]; Uitl. Kapellen 3 (17-21): 63, pl. 229, f. D, E; TL: Coromandel
Papilio hilaria Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 95, pl. 339, f. A, B; TL: Coromandel
Papilio lalage Herbst, 1792; in Jablonsky, Naturs. Schmett. 5: 163, pl. 106, f. 1-2
Papilio alcmeone Fabricius, 1793; Ent. Syst. 3 (1): 193; TL: Coromandel
Papilio titania Fabricius, 1798; Ent. Syst. (Suppl.): 428
Colias jugurthina Godart, 1819; Encyclopédie Méthodique 9 (1): 96; TL: Bengal; Coromandel
Callidryas endeer Boisduval, 1832; in d'Urville, Voy. Astrolabe (Faune ent. Pacif.) 1: 63, pl. 2, f. 3-4
Callidryas phlegeus Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3): 401; TL: Timor
Catopsilia heera Swinhoe, 1885; Proc. Zool. Soc. Lond. 1885: 140; TL: Poona
Catopsilia crocale f. virosa Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 68 e
Catopsilia crocale f. flavescens Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 162, pl. 69 c
Catopsilia crocale f. latilimbata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163; TL: Java
celebica Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 68 e, 69 a
ostentata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 69 a
micromalayana Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163
Catopsilia crocalina Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 68 d
Catopsilia pomona f. citronella Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163; TL: Lesser Sunda Is
Catopsilia pomona f. nivescens Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 69 b
Catopsilia pomona f. perspicua Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 69 b; TL: Celebes
Catopsilia pomona f. serrata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163; TL: Java, Lesser Sunda Is.
Catopsilia aserrata Fruhstorfer, 1910
Catopsilia pomona f. bidotata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163
Catopsilia pomona f. asserata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163
Catopsilia pomona f. siscia Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163; TL: Lesser Sunda Is.
dinkelmanni Lindsey, 1924²; Ent. Rundsch. 41: 2
Catopsilia pura Lindsey, 1924; Ent. Rundsch. 41: 2; TL: India
Catopsilia pomona f. formosana Sonan, 1930; Zephyrus 2 (3): 171; TL: Formosa
Catopsilia pomona ab. matsuwakii Nomura, 1931; Mushi 4 (1): 13
transiens Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 64
joannisi Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 64
fasciata Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 64
deficiens Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 64
Catopsilia inornata Dufrane, 1947
Catopsilia pomona ab. emblematis Murayama, 1961; Tyô to Ga 11 (4): 55, f. 10, 12; TL: Poli, Formosa

Callidryas catilla; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 837
Callidryas crocale; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 837
Callidryas pomona; Piepers & Snellen, 1909, Rhop. Java [1]: 42, pl. 3, f. 6a-k, pl. 4, f. 1a-f
Catopsilia catilla; Moore, [1881], Lepid. Ceylon 1 (3): 122, pl. 47, f. 3, 3a
Catopsilia crocale; Moore, [1881], Lepid. Ceylon 1 (3): 122, pl. 48, f. 1, 1a-b; Wood-Mason & de Nicéville, 1881, J. Asiat. Soc. Bengal 49 Pt.II (4): 236; [BIR], 445; Kudrna, 1974, Atalanta 5: 98; [BOW]: pl. 157, f. 22; [EBW]
Catopsilia pomona catilla; [iNaturalist]
Catopsilia pomona crocale; [iNaturalist]
Catopsilia pomona hilaria; [iNaturalist]
Catopsilia pomona; [BIR], 446; Kudrna, 1974, Atalanta 5: 98; Holloway & Peters, 1976, J. Nat. Hist. 10: 286; [EBW]; [BOW]: pl. 157, f. 23; [BOR], 173; Dunn & Dunn, 1991, Review Austr. Butts.: (120-140) ; [MRS], 214; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 101; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Don Herbison-Evans]

769x513 (~69Kb) underside Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko	769x513 (~108Kb) underside Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko	871x981 (~146Kb) ♂ underside hailand, Chon Buri Province, Pattaya, Ko Khram Island, Saikawe beach, a side of a road through shrubbery at the coast, 3rd February, 2006, Photo © Oleg Kosterin
600x400 (~64Kb) ♀ underside India: Melghat, 8.10.2008, Photo © Kedar Tokekar

C. p. pomona Moluccas - Solomons

Catopsilia pomona alcmeone; [iNaturalist]
Catopsilia pomona jugurtha; [iNaturalist]
Catopsilia pomona pomona; [MRS]; [BAUR], 161; Winhard, 2000, Butterflies of the world 10: 12, pl. 16, f. 1-2, 4-5; [iNaturalist] ; [Yutaka Inayoshi]
Catopsilia pomona pomona f. catilla; [MRS], 214; [BMP]: 96, pl. 9, f. 13
Catopsilia pomona pomona f. crocale; [MRS], 214; [BAUR], 161; [BMP]: 96, pl. 9, f. 21
Catopsilia pomona pomona f. hilaria; [MRS], 214; [BMP]: 96, pl. 9, f. 11
Catopsilia pomona pomona f. jugurtha; [MRS], 214
Catopsilia pomona pomona f. nivescens; [BMP]: 96, pl. 9, f. 12
Catopsilia pomona f. alcmeone;
Catopsilia pomona f. jugurtha;

814x850 (~169Kb) ♂ underside Cambodia, Kep Province, Kep, a pond bank. 21st April 2010, Photo © Oleg Kosterin

1020x787 (~352Kb) ♂ underside Cambodia, Kep Province, a beach. 22nd April 2010, Photo © Oleg Kosterin

1184x1122 (~230Kb) ♂ underside Cambodia, Ratanakiri Province, Ban Lung, at Kan Seng Lake, 31st May 2013, Photo © Oleg Kosterin

*Catopsilia pomona pomona ♀ f. jugurtha*
1134x1133 (~261Kb) ♀ underside Cambodia, Ratanakiri Province, Ban Lung, at Kan Seng Lake, 6th June 2013 (Catopsilia pomona pomona ♀ f. jugurtha), Photo © Oleg Kosterin

*Catopsilia pomona pomona ♀ f. crocale*
1113x989 (~221Kb) ♀ underside Cambodia, Ratanakiri Province, Ban Lung, at Kan Seng Lake, 31st May 2013 (Catopsilia pomona pomona ♀ f. crocale), Photo © Oleg Kosterin

C. p. flava (Butler, 1869) Sulawesi, Salayar, Tanahjampea, Kalao, Kabaena, Muna, Buton, Tukangbesi, Banggai

Callidryas flava Butler, 1869; Ann. Mag. nat. Hist. (4) 4 (21): 202; TL: Celebes

Catopsilia pomona flava; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 101; [iNaturalist]

C. p. rivalis Fruhstorfr, 1910² Sula, New Guinea, Solomon Is., Vanuatu, New Caledona, Australia

Catopsilia crocale flava f. rivalis Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 68; TL: Moluccas and Papua

Catopsilia crocale rivalis; Rothschild, 1915, Novit. Zool. 22 (1): 113
Catopsilia pomona rivalis; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 101

C. p. dionysiades Fruhstorfer, 1911 Talaud, Sangihe

Catopsilia crocale dionysiades Fruhstorfer, 1911; Ent. Rundschau 28 (24): 186; TL: Sangir I.

Catopsilia pomona dionysiades; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 101

Catopsilia pyranthe (Linnaeus, 1758)

Ceylon, India, Burma, Australia. See [maps] Mottled Emigrant

Papilio pyranthe Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 469; TL: Canton, China

Catopsilia pyranthe; Moore, [1881], Lepid. Ceylon 1 (3): 124, pl. 47, f. 2, 2a; Kudrna, 1974, Atalanta 5: 98; [EBW]; [BOR], 170; Dunn & Dunn, 1991, Review Austr. Butts.: (120-140) ; [MRS], 215; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 100; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Don Herbison-Evans]

C. p. pyranthe

Papilio alcyone Cramer, [1775]; Uitl. Kapellen 1 (1-7): 89, pl. 58, f. A,B,C
Papilio gnoma Fabricius, 1775; Syst. Ent.: 808 [dry season form]
Papilio flavocinerascens Goeze, 1779; Ent. Beyträge 3 (1): 183
Papilio philippina Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 139, pl. 361, f. C,D
Papilio nephte Fabricius, 1793; Ent. Syst. 3 (1): 190; TL: China
Papilio ilea Fabricius, 1798; Ent. Syst. (Suppl.): 426 or 421
Callidryas thisorella Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 609; TL: Bengal, India
Catopsilia ab. flavescens Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 63
Catopsilia ab. juncta Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 63
Catopsilia f. transiens Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 63

Catopsilia florella gnoma; [BOR], 170
Catopsilia gnoma; Moore, [1881], Lepid. Ceylon 1 (3): 123, pl. 48, f. 2, 2a
Catopsilia ilea; Moore, [1881], Lepid. Ceylon 1 (3): 124, pl. 47, f. 1, 1a-b
Catopsilia pyranthe pyranthe; [MRS]; [BOR], 170; [BMP]: 95, pl. 9, f. 22; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 7; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 101; [iNaturalist] ; [Yutaka Inayoshi]

1127x866 (~98Kb) ♀ underside Thailand, Chon Buri Province, Pattaya, Jomtien beach, a wasting land behind Jomtien Holiday Hotel, 2nd January 2006, Photo © Oleg Kosterin

1148x1163 (~142Kb) ♀ underside Thailand, Chon Buri Province, Pattaya, Jomtien beach, a wasting land behind Jomtien Holiday Hotel, 2nd January 2006 (ovipositing), Photo © Oleg Kosterin

1068x869 (~93Kb) ♀ underside Cambodia, Siem Reap, shrubs at a channel at the artificial lake of Barai. 8th January 2006, Photo © Oleg Kosterin

C. p. minna (Herbst, 1792) Ceylon, Andamans

Papilio minna Herbst, 1792; in Jablonsky, Naturs. Schmett. 5: pl. 89, f. 1-2
?Mancipium minna Hübner, [1810]; Samml. exot. Schmett. 1: pl. [144]

Callidryas minna; Boisduval, 1836, Hist. nat. Ins., Spec. gén. Lépid. 1: 612
Catopsilia pyranthe minna; [BOR], 170

C. p. chryseis (Drury, [1773])

Papilio chryseis Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1: pl. 12, f. 3-4

Callidryas chryseis; Piepers & Snellen, 1909, Rhop. Java [1]: 39
Catopsilia chryseis; Moore, [1881], Lepid. Ceylon 1 (3): 125, pl. 48, f. 3, 3a
Catopsilia pyranthe chryseis; [MRS], 215

C. p. crokera (Macleay, 1826) NE.Australia, Timor

Pontia crokera Macleay, 1826

Catopsilia pyranthe crokera; [BAUR], 161
crookera[sic, crokera]; Wallace, 1867

C. p. lactea (Butler, 1870)

Callidryas lactea Butler, 1870; Ann. Mag. nat. Hist. (4) 5 (29): 361; TL: South-Sea Islands

Catopsilia pyranthe lactea; Holloway & Peters, 1976, J. Nat. Hist. 10: 285

C. p. evangelina (Butler, 1870)

Callidryas evangelina Butler, 1870; Trans. ent. Soc. Lond. 1870 (1): 11; TL: Flores

Catopsilia pyranthe evangelina; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 8

C. p. pythias Waterhouse & Lyell, 1914 N.Australia, New South Wales

pyranthe pythias Waterhouse & Lyell, 1914; Butts Australia: 149, f. 509
lacteola Waterhouse & Lyell, 1914; Butts Australia: 150, f. 510-511

Catopsilia scylla (Linnaeus, 1763)

Siam, Malayasia, Moluccas. See [maps] Orange Emigrant Yellow Migrant

Papilio scylla Linnaeus, 1763; Amoenitates Acad. 6: 404
Papilio scylla Linnaeus, 1764; Mus. Lud. Ulr.: 242; TL: Java
Catopsilia scylla ab. ascylla Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2): 275; TL: W.Java; Lombok; Sumbawa
Callidryas scylla ♀ f. obscura van den Bergh, 1917; Tijdschr. Ent. 60: 268

Callidryas scylla; Piepers & Snellen, 1909, Rhop. Java [1]: 40, pl. 3, f. 5a-b
Catopsilia scylla; [EBW]; [EOB]; [BOW]: pl. 157, f. 21, pl. 158, f. 1; [BOR], 173; [MRS], 215; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 100; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Don Herbison-Evans]

C. s. scylla Java, Bali, Lombok, Bawean

Catopsilia scylla scylla; [BOR], 173

C. s. cornelia (Fabricius, 1787) S.Burma, Peninsular Malaya, Singapore, Sumatra

Papilio cornelia Fabricius, 1787; Mantissa Insectorum 2: 21, no. 229

Catopsilia scylla cornelia; [BOR], 173; [BMP]: 98, pl. 9, f. 10; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 10; [iNaturalist] ; [Yutaka Inayoshi]

700x903 (~73Kb) ♂ underside Thailand, Chon Buri Province, Pattaya environs, Ko Lan island, ruderal vegeation at a roadside. 27th January 2005, Photo © Oleg Kosterin

804x1086 (~106Kb) ♂ underside Thailand, Chon Buri Province, Pattaya, Ko Khram Island, Saikawe beach, a side of a road through shrubbery at the coast, 3rd February, 2006, Photo © Oleg Kosterin

1173x1125 (~137Kb) ♂ underside Thailand, Chon Buri Province, Pattaya, a small wood at a grassy swamp at the Jomtien beach, between B.O. Guesthouse and Jomtien Metro Condotel, 2nd January, 2006, Photo © Oleg Kosterin

C. s. etesia (Hewitson, 1867) Northern Territory - Cape York

Callidryas etesia Hewitson, 1867; Ill. exot. Butts [1] (Callidryas and Eronia): [63], pl. [33], f. 5-6; TL: Australia (Queensland)

Catopsilia scylla etesia; [BAUR], 161

C. s. asema Staudinger, 1885² Philippines, Sulawesi, Salayar, Kalao, Kabaena, Buton, Tukangbesi

scylla asema Staudinger, 1885²; in Staudinger & Schatz, Exot. Schmett. 1 (7): 39, (5): pl. 21
minacia Fruhstorfer, 1910²; in Seitz, Gross-Schmett. Erde 9: 163, pl. 68 b

Catopsilia scylla asema; [BOR], 173; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 100; [iNaturalist]

C. s. scylloides Fruhstorfer, 1903 Babber

Catopsilia scylla scylloides Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2): 275; TL: Babber

C. s. bangkejana Fruhstorfer, 1903 Banggai I., Sula I.

Catopsilia scylla bangkejana Fruhstorfer, 1903; Soc. Ent. 18 (10): 73; TL: Bangkei I.

Catopsilia scylla bangkejana; [BOR], 173; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 100

C. s. sidra Fruhstorfer, 1910 Sumbawa

Catopsilia scylla sidra Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 163, pl. 68 b-c

Catopsilia scylla sidra; [BOR], 173

C. s. praerubida Fruhstorfer, 1910 Flores

Catopsilia scylla praerubida Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 164

Catopsilia scylla praerubida; [BOR], 173

C. s. moluccarum Fruhstorfer, 1910 Moluccas

Catopsilia scylla moluccarum Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9: 165, pl. 68 row c; TL: Moluccas

Catopsilia scylla moluccarum; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 11-12

C. s. philea Krüger, 1929 Sumba

scylla philea Krüger, 1929; Int. ent. Zs. 23 (4): 58; TL: Sumba

C. s. sumbana Krüger, 1929 Sumba

scylla sumbana Krüger, 1929; Int. ent. Zs. 23 (4): 58; TL: Sumba
rosea Krüger, 1929; Int. ent. Zs. 23 (4): 59

Catopsilia gorgophone (Boisduval, 1836)

Australia (Cairns - Sydney). See [maps] Yellow Migrant

Callidryas gorgophone Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 632; TL: Queensland
Callidryas hinda Butler, 1870; Lepid. Exotica (4): 31, pl. 12, f. 9-10

Catopsilia gorgophone; Dunn & Dunn, 1991, Review Austr. Butts.: (120-140) ; [CoL] ; [iNaturalist] ; [Don Herbison-Evans]
Catopsilia hinda; [iNaturalist]
Catopsilia scylla gorgophone; [BAUR], 161; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 13

C. g. novacaledonia Viette, 1950 New Caledonia

Catopsilia gorgophone novacaledonia Viette, 1950; Fn. Emp. franc. 13: 40

Catopsilia scylla novacaledonia; Holloway & Peters, 1976, J. Nat. Hist. 10: 286

Catopsilia florella (Fabricius, 1775)

Africa, Madagascar, Canary Islands, Ceylon, India, Burma, China. See [maps] African Migrant

Papilio florella Fabricius, 1775; Syst. Ent.: 479; TL: Sierra Leone
Colias pyrene Swainson, 1821; Zool. Illustr. (1) 1: pl. 51
- = ; [AfrL]
Callidryas hyblaea Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 612; TL: Comoro Is.
- = ; [AfrL]
Callidryas rhadia Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 617
- = ; [AfrL]
Pontia marcellina Bertoloni, 1850; Mem. Accad. Bologna 2: 178
- = ; [AfrL]
Catopsilia aleurona Butler, 1876; Ann. Mag. nat. Hist. (4) 18 (108): 489
- = ; [AfrL]
Catopsilia rufosparsa Butler, 1880; Ann. Mag. nat. Hist. (5) 5 (29): 395; TL: Madagascar
- = ; [AfrL]
Callidryas swainsoni Westwood, 1881; in Oates, Matabeland: 335
- = ; [AfrL]
Catopsilia florella ab. subpyrene Strand, 1911; Int. ent. Zs. 4 (41) (41): 220; TL: Tanzania
- = ; [AfrL]
?ornata Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 62
Catopsilia florella florella f. inornata Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 62
- = ; [AfrL]
Catopsilia florella florella ab. biannulata Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 62
- = ; [AfrL]
Catopsilia florella florella ab. houzeaui Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2): 63
- = ; [AfrL]
Catopsilia florella ♀-f. peregrina Stoneham, 1957; Bull. Stoneham Mus. (70): [2]; TL: Kenya
- = ; [AfrL]
Catopsilia florella ♀-f. duplicata Stoneham, 1957; Bull. Stoneham Mus. (70): [2]; TL: Kenya
- = ; [AfrL]
Catopsilia florella (f) wandriana Stoneham, 1957; Bull. Stoneham Mus. (70): [2]; TL: Kenya
- = ; [AfrL]

Catopsilia florella; [EBW]; [H&R]; [BMAT]: 18, pl. 6, f. 34-36; [BAFR], 53; [PBSA]; [BK]; [MadL]; [AfrL]; [Otakar Kudrna]; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE]
Catopsilia florella f. hyblaea; [BAFR], 52, fig. ♀ form.)
Catopsilia florella f. pyrene; [BAFR], 52, fig. ♀ form.)

1055x929 (~382Kb) underside Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin

C. f. florella

Catopsilia florella florella; [BOR], 170; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 6, 9

Catopsilia thauruma (Reakirt, 1866)

Madagascar, Mauritius. See [maps]

Callidryas thauruma Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3): 238; TL: Madagascar
Callidryas fiaduna Hewitson, 1867; Ill. exot. Butts [1] (Callidryas and Eronia): [63], pl. [33], f. 1-4; TL: Madagascar
- = ; [AfrL]
Eronia grandidieri Mabille, 1877; Bull. Soc. ent. Fr. (5) 7: xxxviii; TL: Madagascar
- = ; [AfrL]
Catopsilia decipiens Butler, 1880; Ann. Mag. nat. Hist. (5) 5 (28): 338; TL: Madagascar
- = ; [AfrL]
Catopsilia grandidieri var. mauritiana Le Cerf, 1916; Bull. Soc. Ent. Fr. 1916: 112
- = ; [AfrL]
Catopsilia mabillei Neustetter, 1929; Int. ent. Zs. 23: 336; TL: Madagascar
- = ; [MadL]: fn. #15; [AfrL]

Catopsilia grandidieri; [EBW]; [CoL]
Catopsilia mabillei; [EBW]; [CoL]
Catopsilia thauruma; [EBW]; [BAFR], 53; Winhard, 2000, Butterflies of the world 10: 13, pl. 16, f. 3; [MadL]; [AfrL]; [CoL] ; [iNaturalist]
Catopsilia thauruma f. grandidieri; [BAFR], 53, fig.

References:

[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[Don Herbison-Evans] and Stella A. Crossley
Australian Caterpillars and their Butterflies and Moths
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Kedar Tokekar]
India
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Olli Vesikko]
http://koti.welho.com/ovesikko/
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[²] This may require parentheses or not (original genus not verified)

Some related literature:

van den Bergh, 1917
Callidryas scylla L. ♀. Nieuwe vorm: Obscura van den Bergh Tijdschr. Ent. 60 : 268-269, pl. 3
Bertoloni, 1850
Illustrazione dei prodotti naturali del Mozambico. Dissertazione IV. Insetti Lepidotteri Diurni Mem. Accad. Bologna 2: : 165-188
Boisduval, 1832
Voyage de découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les années 1826-1827-1828-1829, sous le commandément de M. J. Dumont d'Urville. Faune entomologique de l'Océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Lépidoptères in d'Urville, Voy. Astrolabe (Faune ent. Pacif.) 1 : 1-267, pl. 1-5
Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, 1870
Lepidoptera Exotica, or descriptions and illustrations of exotic Lepidoptera Lepid. Exotica (1): 1-8, pl. 1-3 (1869), (2): 9-16, pl. 4-6 (1869), (3): 17-24, pl. 7-9 (1869), (4): 25-32, pl. 10-12 (1870), (5): 33-42, pl. 13-15 (1870), (6): 43-50, pl. 16-19 (1870), (7): 51-58, pl. 20-22 (1870), (8): 59-66, pl. 23-25 (1871), (9): 67-74, pl. 26-28 (1871), (10): 75-84, pl. 29-32 (1871), (11): 85-94, pl. 33-35 (1872), (12): 95-104, pl. 36-38 (1872), (13): 105-114, pl. 39-41 (1872), (14): [115],[116],117-120, pl. 42-44 (1872), (15): 121-128, pl. 45-48 (1873), (17): 137-144, pl. 49-51 (1873), (18): 145-152, pl. 52-54 (1873), (19): 153-162, pl. 55-57 (1873), (20): 163-174, pl. 58-60 (1874), (21): 175-190, pl. 61-64 (1874)
Butler, 1869
Descriptions of three new Species of Callidryas Ann. Mag. nat. Hist. (4) 4 (21) : 202-203
Butler, 1870
List of Species in a small Collection of Butterflies from the South Seas Ann. Mag. nat. Hist. (4) 5 (29) : 357-361
Butler, 1870
Descriptions of six new species of Callidryas Trans. ent. Soc. Lond. 1870 (1) : 9-12
Butler, 1876
On a Collection of Lepidoptera recently received from Abyssinia Ann. Mag. nat. Hist. (4) 18 (108) : 480-490
Butler, 1880
On a collection of Lepidoptera from Madagascar, with descriptions of new Genera and Species Ann. Mag. nat. Hist. (5) 5 (28): 333-344, (29): 384-395
Le Cerf, 1916
Note sur deux Pierides [Lep. Rhopal.] de l'ile Maurice et de l'ilot d'Astove Bull. Soc. Ent. Fr. 1916 : 112-113
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
[BMP]; Corbet & Pendlebury, 1992
The butterflies of the Malay Peninsula. Fourth Edition revised by J. N. Eliot with plates by Bernard D'Abrera Butterflies Malay Peninsula (4th edn) : 1-595, pl. 1-69
Cramer, [1775]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
Cramer, [1779]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 3 (17-21): 1-104, pl. 193-252 (1779), (22): 105-128, pl. 253-264 ([1780]), (23-24): 129-176, pl. 265-288 (1780)
[BAUR]; D'Abrera, 1977
Butterflies of the Australian Region (2nd edn) Butts. Aust. Region: 1-416
[BAFR]; D'Abrera, 1980
Butterflies of the Afrotropical Region: -
[BOR]; D'Abrera, 1982-1986
Butterflies of the Oriental Region I-III
[PBSA]; Dickson & Kroon, 1978
Pennington's Butterflies of Southern Africa
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1 : pl. 1-50 (1770), (1773) , : 1-123 (1837)
Dufrane, 1947
Pieridae Bull. Ann. Soc. ent. Belg. 83 (1/2) : 46-73
Dunn & Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy Review Austr. Butts. (1): 1-196, (2): 197-335, (3): 336-512, (4): 513-660
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus Syst. Ent. : 1-832
Fabricius, 1787
Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus Mantissa Insectorum 2 : 1-382
Fabricius, 1793
Entomologia Systematica emendata et aucta. Ent. Syst. 3 (1) : 1-487
Fabricius, 1798
Supplementum Entomologiae Systematicae Ent. Syst. (Suppl.) : 1-572, (index) 1-53
[EOB]; Feltwell, 1993
The Encyclopedia of Butterflies
Fruhstorfer, 1903
Verzeichniss der in Tonkin, Annam und Siam gesammelten Pieriden und Besprechung verwandter Formen Dt. ent. Z. Iris 15 (2) : 269-305 (1 May)
Fruhstorfer, 1903
Neue Rhopaloceren des Malayischen Archipels Soc. Ent. 18 (10) : 73 (15 August)
Fruhstorfer, 1910
2. Familie: Pieridae, Weisslinge in Seitz, Gross-Schmett. Erde 9 : 119-120 (1909), : 121-144 (14 January, 1910), : 145-152 (11 February, 1910), : 153-168 (2 April, 1910), : 169-176 (4 April, 1910), : 177-190 (20 May, 1910)
Fruhstorfer, 1911
Neue Pieriden aus der Sammlung Staudinger des Berliner Zoologischen Museums Ent. Rundschau 28 (24) : 185-187 (16 December)
Godart, 1819
Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Goeze, 1779
Entomologische Beyträge zu des Ritter Linné zwölften Ausgabe des Natursystems Ent. Beyträge 3 (1) : 1-390
Herbst, 1792
Natursystem aller bekannten in- und auslandischen Insekten als eine Fortsetzun der von Büffonschen Naturgeschichte in Jablonsky, Naturs. Schmett. 5 : 1-231
Hewitson, 1867
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson. Ill. exot. Butts [1] : (Systematic Index) I, (Ornithoptera and Papilio): [1-2], pl. [1] (1855), (Papilio II): [3-4], pl. [2] (1856), (Papilio III): [5-6], pl. [3] (1858), (Papilio IV): [7-8], pl. [4] (1859), (Papilio V): [9-10], pl. [5] (1861), (Papilio VI): [11-12], pl. [6] (1864), (Papilio VII-VIII): [13-16], pl. [7-8] (1865), (Papilio IX): [17-18], pl. [9] (1868), (Papilio X-XII): [19-22], pl. [10-12] (1869), (Papilio XIII): [23-24], pl. [13] (1873), (Papilio XIV): [25-26], pl. [14] (1875), (Papilio XV): [27-28], pl. [15] (1877), (Pieris I): [29-30], pl. [16] (1852), (Pieris II): [31-32], pl. [17] (1853), (Pieris III-VI): [33-40], pl. [18-21] (1861), (Pieris VII): [41-42], pl. [22] (1862), (Pieris VIII): [43-44], pl. [23] (1866), (Euterpe): [45-46], pl. [24] (1860), (Euterpe II): [47-48], pl. [25] (1872), (Euterpe and Leptalis): [49-50], pl. [26] (1853), (Leptalis II): [51-52], pl. [27] (1857), (Leptalis III): [53-54], pl. [28] (1858), (Leptalis IV): [55-56], pl. [29] (1860), (Leptalis V-VII): [57-62], pl. [30-32] (1870), (Callidryas and Eronia): [63-64], pl. [33] (1867), (Euploea): [65-66], pl. [34] (1858), (Euploea II): [67-68], pl. [35] (1866), (Heliconia I): [69-70], pl. [36] (1853), (Heliconia II-III): [71-74], pl. [37-38] (1854), (Heliconia IV): [75-76], pl. [39] (1858), (Heliconia V): [77-78], pl. [40] (1867), (Heliconia VI): [79-80], pl. [41] (1871), (Heliconia and Tithorea VII): [81-82], pl. [42] (1873), (Heliconia VIII): [83-84], pl. [43] (1875), (Athesis): [85-86], pl. [44] (1872), (Mechanitis I): [87-88], pl. [45] (1856), (Mechanitis II-III): [89-92], pl. [46-47] (1860), (Ithomia I): [93-94], pl. [48] (1851), (Ithomia II): [95-96], pl. [49] (1852), (Ithomia III): [97-98], pl. [50] (1853), (Ithomia IV-V): [99-102], pl. [51-52] (1854), (Ithomia VI-XIII): [103-118], pl. [53-60] (1855)
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
Holloway & Peters, 1976
The butterflies of New Caledonia and the Loyalty Islands J. Nat. Hist. 10 : 273-318
Hübner, [1810]
Sammlung exotischer Schmetterlinge, Vol. 1 (1806 - [1819]) Samml. exot. Schmett. 1 : [1-4] (1807), : [5-12] (1807-1809), : [13-36] (1823-1824), : pl. [1-2], [5-8], [12], [15], [29], [47] (1806), : pl. [9], [11], [17], [21], [25-27], [32], [35], [43-44], [56], [60], [79], [102], [104], [116], [141], [148], [150-151], [155], [159], [200] (1807), : pl. [19], [39], [57], [73], [82], [114], [157], [167], [170], [173], [187] (1807), : pl. [3], [31], [37], [46], [54], [58], [62], [76], [84], [92], [95], [105], [112-113], [117], [137-138], [146], [154], [158], [180], [209] (1808), : pl. [16], [28], [65], [89], [142], [160], [162], [166], [182], [188], [193] (1809), : pl. [4], [22], [42], [53], [61], [88], [90], [97], [100], [107], [110], [118], [120], [140], [143], [149], [161], [171], [178], [184], [196], [202] (1809), : pl. [10], [18], [30], [41], [51], [80], [144], [172], [189] (1810), : pl. [36], [70], [87], [91], [101], [124], [129], [177], [183], [207], [210] (1811), : pl. [20], [38], [45], [55], [85], [98], [103], [108], [136], [139], [152-153], [156], [163], [179], [186], [190], [205-206] (1812), : pl. [13-14], [40], [48], [50], [67], [74], [86], [119], [121], [130], [135], [175-176], [197-199], [204], [212] (1813), : pl. [72], [75], [77-78], [83], [93], [109], [111], [115], [128], [131], [165], [174], [194] (1813), : pl. [24], [49], [52], [63-64], [66], [68-69], [71], [81], [123], [145], [181], [192] (1814), : pl. [126], [132-134], [168] (1815), : pl. [23], [96], [122], [125], [169], [195] (1816), : pl. [33], [99], [127], [187], [185], [213] (1817), : pl. [59], [106], [147], [164], [208], [211] (1818), : pl. [34], [94], [191], [201] (1819), : (Index Systematicus) [1-4] (1819 - April 1823)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Krüger, 1929
Neue tropische Falter Int. ent. Zs. 23 (4) : 58-59
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
[MadL]; Lees, Kremen & Raharitsimba, 2003
Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist in Goodman & Benstead, Nat. Hist. Madagascar : 762-793
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1763
in Johansson (Thesis), Centuria Insectorum Amoenitates Acad. 6 : 384-415
Linnaeus, 1764
Museum S'ae R'ae M'tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque Mus. Lud. Ulr. : 1-720
Mabille, 1877
[no title] Bull. Soc. ent. Fr. (5) 7 : xxxvii-xxxix [37-39], : xxxix-xl [39-40], : lxxi-lxxiii [71-73]
Moore, 1878
A list of the Lepidopterous Insects collected by Mr. Ossian Limborg in Upper Tenasserim, with Descriptions of new Species Proc. zool. Soc. Lond. 1878 (4) : 821-859, pl. 51-53 (1879)
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Murayama, 1961
An unrecorded and some aberrant butterflies from Formosa [in Japanese] Tyô to Ga 11 (4): 54-61
Neustetter, 1929
Eine neue Catopsilia von Madagascar Int. ent. Zs. 23 : 336-337
Nomura, 1931
Some new aberrant Forms of the Formosan Butterflies Mushi 4 (1) : 10-14
Piepers & Snellen, 1909
The Rhopalocera of Java. Pieridae Rhop. Java [1] : 1-65, pl. 1-4
Reakirt, 1866
Descriptions of some new species of Diurnal Lepidoptera Proc. Acad. nat. Sci. Philad. 18 (3) : 238-249
Rothschild, 1915
On Lepidoptera from the islands of Ceram (Seran), Buru, Bali, and Misol Novit. Zool. 22 (1): 105-144, (2): 209-227
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Sonan, 1930
Notes on some butterflies from Formosa [in Japanese] Zephyrus 2 (3) : 165-176
Staudinger, 1885
Exotische Tagfalter in sysmatischer Reihenfolge mit Berücksichtigung neuer Arten in Staudinger & Schatz, Exot. Schmett. 1 (1): 3-6, pl. 1-5 (1884), (2): 7-10, pl. 6-10 (1884), (3): 11-14, pl. 11-15 (1884), (4): 15-22, pl. 16-20 (1884), (5): 23-30, pl. 21-25 (1884), (6): 31-38, pl. 26-30 (1884), (7): 39-46, pl. 31-35 (1885), (8): 47-58, pl. 30-40 (1885), (9): 59-82, pl. 41-50 (1885), (10): 83-102, pl. 51-60 (1885), (11): 103-122, pl. 61-65 (1886), (12): 123-138, pl. 66-70 (1886), (13): 139-158, pl. 71-75 (1886), (14): 159-174, pl. 76-80 (1886), (15): 175-194, pl. 81-85 (1887), (16): 195-214, pl. 86-90 (1887), (17): 215-234, pl. 91-95 (1887), (18): 235-270, pl. 96-98 (1888), (19): 1-2,271-333, pl. 99-100 (1888)
Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Stoneham, 1957
Two new generic names, and new subspecies and forms of Pieridae Bull. Stoneham Mus. (70) : 1-3
Strand, 1911
Neue afrikanische Pierididen un Nymphaliden Int. ent. Zs. 4 (41): (41): 219-220, (42): 226-227
Swainson, 1821
Zoological illustrations, or original figures and descriptions of new, rare, or interesting animals, selected chiefly from the classes of ornithology, entomology, and conchology, and arranged on the principles of Cuvier and other modern zoologists Zool. Illustr. (1) 1: pl. 1-18 (1820), 1: pl. 19-66 (1821), 2: pl. 67-83 (1821), 2: pl. 84-119 (1822), 3: pl. 120-134 (1822), 3: pl. 135-182 (1823)
Swinhoe, 1885
On the Lepidoptera of Bombay and the Deccan. Part I-IV Proc. Zool. Soc. Lond. 1885 : 124-148, pl. 9 (I. Rhopalocera), : 287-307, pl. 20-21 (II. Heterocera), : 447-476, pl. 27-28 (III. Heterocera), : 852-886, pl. 56-57 (IV. Heterocera)
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Wallace, 1867
On the Pieridae of the Indian and Australian Regions Trans. ent. Soc. Lond. (3) 4 (3) : 301-416, pl. 6-9
Waterhouse & Lyell, 1914
The Butterflies of Australia, a Monograph of the Australian Rhopalocera, introducing ... Butts Australia : 1-239, pl. 1-42
Westwood, 1881
Entomology in, Matabele Land and the Victoria Falls in Oates, Matabeland : 331-365, pl. E-H
Winhard, 2000
Pieridae I Butterflies of the world 10 : 1-40, pl. title,1-48,back
Wood-Mason & de Nicéville, 1881
List of Diurnal Lepidoptera from Port Blair, Andaman Islands J. Asiat. Soc. Bengal 49 Pt.II (4) : 223-243, pl. 13
[BIR]; Wynter-Blyth, 1957
Butterflies of the Indian Region (1982 Reprint)
Zhang, Cong, Shen, Opler & Grishin, 2021
Genomics-guided refinement of butterfly taxonomy Taxonomic Rep. 9 (3) : 1-55

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