Pyrginae  Burmeister, 1878
  Dicot Skippers 
  Grizzled Skippers 
  Spread-wing Skippers 
  Flats 
 
Achalarus — 5 spp. 
Aguna — 26 spp. 
Astraptes — 30 spp. 
Augiades — 2 spp. 
Aurina — 1 sp. 
Autochton — 13 spp. 
Bungalotis — 10 spp. 
Cabares — 2 spp. 
Cabirus — 1 sp. 
Calliades — 2 spp. 
Celaenorrhinus 
  Flats 
 
 — 119 spp. 
Cephise — 12 spp. 
Chioides — 10 spp. 
Chrysoplectrum — 10 spp. 
Codatractus — 15 spp. 
Drephalys — 18 spp. 
Dyscophellus — 9 spp. 
Entheus — 14 spp. 
Epargyreus — 19 spp. 
Heronia — 1 sp. 
Hyalothyrus — 5 spp. 
Hypocryptothrix — 1 sp. 
Narcosius — 12 spp. 
Nascus — 6 spp. 
Ocyba — 1 sp. 
Oileides — 4 spp. 
Phanus — 9 spp. 
Phareas — 1 sp. 
Phocides — 21 spp. 
Polygonus — 2 spp. 
Polythrix — 14 spp. 
Porphyrogenes — 14 spp. 
Proteides — 2 spp. 
Ridens — 20 spp. 
Salatis — 7 spp. 
Sarmientola — 8 spp. 
Tarsoctenus — 3 spp. 
Thessia — 2 spp. 
Thorybes — 7 spp. 
Typhedanus — 11 spp. 
Udranomia — 4 spp. 
Urbanus — 36 spp. 
Venada — 5 spp. 
Zestusa — 4 spp. 
 
[NL4A]  places new world genera into Eudamini Celaenorhini  go there also?
Tribe Pyrgini  
Pyrgini  (Pyrginae); [NL4A] , 42
 
  
In addition to these, some other genera may belong... -- msa
Muschampia — 19 spp. 
Spialia 
  Grizzled Skippers, Sandmen 
 
 — 28 spp. 
Gomalia — 1 sp. 
 
Achlyodes — 5 spp. 
Aethilla — 10 spp. 
Anastrus — 11 spp. 
Anisochoria — 10 spp. 
Antigonus — 9 spp. 
Arteurotia — 1 sp. 
Atarnes — 2 spp. 
Bolla — 28 spp. 
Burca — 5 spp. 
Camptopleura — 6 spp. 
Carrhenes — 13 spp. 
Celotes — 2 spp. 
Charidia — 2 spp. 
Chiomara — 5 spp. 
Clito — 11 spp. 
Cogia — 15 spp. 
Conognathus — 1 sp. 
Cornuphallus — 3 spp. 
Cycloglypha — 6 spp. 
Cyclosemia — 9 spp. 
Diaeus — 4 spp. 
Doberes — 2 spp. 
Ebrietas — 7 spp. 
Ectomis — 1 sp. 
Ephyriades — 4 spp. 
Eracon — 6 spp. 
Erynnis 
  Duskywings 
 
 — 24 spp. 
Gesta — 5 spp. 
Gindanes — 3 spp. 
Gorgopas — 5 spp. 
Gorgythion — 6 spp. 
Grais — 1 sp. 
Haemactis — 3 spp. 
Helias — 3 spp. 
Heliopetes — 14 spp. 
Heliopyrgus — 3 spp. 
Hesperopsis — 3 spp. 
Iliana — 4 spp. 
Jera — 1 sp. 
Marela — 2 spp. 
Mictris — 1 sp. 
Milanion — 6 spp. 
Mimia — 2 spp. 
Morvina — 3 spp. 
Mylon — 15 spp. 
Myrinia — 6 spp. 
Nerula — 2 spp. 
Nisoniades — 34 spp. 
Noctuana — 6 spp. 
Ocella — 3 spp. 
Oechydrus — 1 sp. 
Onenses — 2 spp. 
Ouleus — 11 spp. 
Paches — 6 spp. 
Pachyneuria — 11 spp. 
Paracogia — 1 sp. 
Paramimus — 2 spp. 
Pellicia — 16 spp. 
Pholisora — 3 spp. 
Plumbago — 3 spp. 
Polyctor — 6 spp. 
Potamanaxas — 16 spp. 
Pseudodrephalys — 3 spp. 
Pyrdalus — 1 sp. 
Pyrgus 
  Grizzled Skippers 
  Checkered Skippers 
 
 — 48 spp. 
 4 spp. 
Pythonides — 20 spp. 
Quadrus — 11 spp. 
Sophista — 2 spp. 
Sostrata — 9 spp. 
Spathilepia — 1 sp. 
Speculum — 1 sp. 
Spioniades — 3 spp. 
Staphylus — 55 spp. 
Systasea — 3 spp. 
Telemiades — 18 spp. 
Theagenes — 3 spp. 
Timochares — 3 spp. 
Timochreon — 2 spp. 
Tosta — 7 spp. 
Trina — 1 sp. 
Viola — 6 spp. 
Windia — 1 sp. 
Xenophanes — 1 sp. 
Xispia — 2 spp. 
Zera — 8 spp. 
Zobera — 3 spp. 
Zopyrion — 5 spp. 
 
Eretis 
  Elfs, Small Flats 
 
 — 11 spp. 
Eagris — 9 spp. 
Netrobalane 
  Buff-tipped Skippers 
 
 — 1 sp. 
 
In addition to these, some other genera may belong... -- msa
Tapena — 1 sp. 
Darpa — 4 spp. 
Odina — 2 spp. 
Sarangesa 
  Small Flats, Elfins 
 
 — 26 spp. 
Pseudocoladenia — 5 spp. 
Coladenia 
  Pied Flats 
 
 — 20 spp. 
Satarupa — 7 spp. 
Seseria — 6 spp. 
Pintara — 5 spp. 
Chamunda — 1 sp. 
Gerosis — 8 spp. 
Mooreana — 3 spp. 
Tagiades 
  Clouded Flats 
  Water Flats 
  Snow Flats 
 
 — 18 spp. 
Odontoptilum — 3 spp. 
Abraximorpha — 4 spp. 
Caprona 
  Ragged Skippers 
 
 — 6 spp. 
Leucochitonea 
  White Cloaked Skippers 
 
 — 3 spp. 
Ctenoptilum — 2 spp. 
Daimio — 1 sp. 
 
Abantis 
  Paradise Skippers 
 
 — 23 spp. 
Euschemon — 1 sp. 
 
  
1420x1135 (~358Kb) upperside  Ethiopia, Oromia, Debre Zeyit, Lake Bishoftu bank. 10th August 2012, Photo © Oleg Kosterin 
The exact identification of this species is still unknown,
but tentatively assumed to belong into this group. 
2.3.2022 (1425)
References: 
[CoL] Catalogue of Lifehttps://doi.org/10.48580/dfz8d 
 [EVERSMANNIA] Eversmannia contents 
 [Oleg Kosterin] Institue of Cytology & Geneticse-mail:  Oleg Kosterin ;  http://pisum.bionet.nsc.ru/kosterin/ 
 [iNaturalist] iNaturalist 
  Some related literature: 
Korb & Bolshakov, 2011Eversmannia Suppl.  2  [NL4A]; Lamas, 2004Atlas of Neotropical Lepidoptera 
 [BK]; Larsen, 1996Butterflies of Kenya 
 [RFEL]; Leley, 2016Cat. ins. Russian Far East.  2   If you have corrections, comments or
 information to add into these pages, just send mail to
Markku Savela