Theclinae  
  Nopsasiivet 
  Hairstreaks 
  Strong Blues 
 
Aphnaeinae  (Lycaenidae); Clench, 1965, Mem. Amer. Ent. Soc.  19  : 357 
Theclinae ;  [CoL ]   ;  [iNaturalist ]   
Theclinae  (Lycaenidae); Clench, 1965, Mem. Amer. Ent. Soc.  19  : 331 ; Eliot, 1973, Bull. Br. Mus. nat. Hist.  (Ent.) 28  (6) : 381 ; Vane-Wright & de Jong, 2003,  Zool. Verh. Leiden  343  : 120; [NL4A] , 118;  [AfrL ]; Korb & Bolshakov, 2011,  Eversmannia Suppl.  2  : 68; [RFEL] 
 
  
 
Artopoetes   Chapman, 1909 
 — 2 spp. 
Laeosopis   Rambur, 1858 
 — 1 sp. 
Thecla   Fabricius, 1807 
 — 3 spp. 
 1 sp. 
Shirozua   Sibatani & Ito, 1942 
 — 2 spp. 
Cordelia   Shirôzu & Yamamoto, 1956 
 — 3 spp. 
Gonerilia   Shirôzu & Yamamoto, 1956 
 — 5 spp. 
Coreana   Tutt, [1907] 
 — 1 sp. 
Ussuriana   Tutt, [1907] 
 — 4 spp. 
Chaetoprocta   de Nicéville in Marshall & de Nicéville, 1890 
 — 3 spp. 
Protantigius   Shirôzu & Yamamoto, 1956 
 — 1 sp. 
Leucantigius   Shirôzu & Murayama, 1951 
 — 1 sp. 
Ravenna   Shirôzu & Yamamoto, 1956 
 — 1 sp. 
Antigius   Sibatani & Ito, 1942 
 — 5 spp. 
Wagimo   Sibatani & Ito, 1942 
 — 6 spp. 
Araragi   Sibatani & Ito, 1942 
 — 3 spp. 
Japonica   Tutt, [1907] 
 — 5 spp. 
Hypaurotis   Scudder, 1876 
 — 1 sp. 
Habrodais   Scudder, 1876 
 — 2 spp. 
Euaspa   Moore, 1884 
 — 12 spp. 
Howarthia   Shirôzu & Yamamoto, 1956 
 — 13 spp. 
Teratozephyrus   Sibatani, 1946 
 — 10 spp. 
Esakiozephyrus   Shirôzu & Yamamoto, 1956 
 — 12 spp. 
Neozephyrus   Sibatani & Ito, 1942 
 — 7 spp. 
Chrysozephyrus   Shirôzu & Yamamoto, 1956 
 — 70 spp. 
Iratsume   Sibatani & Ito, 1942 
 — 1 sp. 
Favonius   Sibatani & Ito, 1942 
 — 14 spp. 
 1 sp. 
Austrozephyrus   Howarth, 1957 
 — 3 spp. 
 
Subtribe Oxylidina  
Oxylidina  (Theclini);  [AfrL ]
Oxylidini  (Theclinae);  
 
  
 
Amblypodia   Horsfield, [1829] 
 — 3 spp. 
Iraota   Moore, [1881] 
  Silverstreak Blues 
 
 — 6 spp. 
Larva  on
Ficus   ,
;  
Myrina   Fabricius, 1807 
  Fig Tree Blues 
 
 — 5 spp. 
 
Hemiolaus   Aurivillius, 1922 
 — 4 spp. 
Hypolycaena   C. & R. Felder, 1862 
  Hairstreaks 
  Tits 
 
 — 46 spp. 
Zeltus   de Nicéville in Marshall & de Nicéville, 1890 
 — 1 sp. 
Chliaria   Moore, 1884 
  Tits 
 
 — 5 spp. 
Leptomyrina   Butler, 1898 
 — 9 spp. 
 
Subtribe Deudorigini   Doherty, 1886 
 
 
Deudorix   Hewitson, 1863 
  Playboys 
  Cornelians 
 
 — 62 spp. 
Artipe   Boisduval, 1870 
 — 4 spp. 
Hypomyrina   Druce, 1891 
 — 4 spp. 
Pilodeudorix   Druce, 1891 
  Playboys 
 
 — 52 spp. 
Sithon   Hübner, [1819] 
 — 2 spp. 
Sinthusa   Moore, 1884 
  Sparks 
 
 — 18 spp. 
Araotes   Doherty, 1889 
 — 2 spp. 
Bindahara   Moore, [1881] 
 — 3 spp. 
Rapala   Moore, [1881] 
  Flashes 
 
 — 55 spp. 
Larva  on
Anidesma  ,
Ziziphus  xylopyrus ,
Lantana camara  ,
Elaeagnus  ferruginea ,
Quisqualis   ,
Acacia caesia  ,
Astilbe rivularis  ,
;  
 
Capys   Hewitson, 1865 
  Protea Butteflies 
 
 — 17 spp. 
Pamela   Hemming, 1935 
 — 1 sp. 
 
Paradeudorix   Libert, 2004 
 — 8 spp. 
Qinorapala   Chou & Wang, 1995 
 
 
Drina   de Nicéville in Marshall & de Nicéville, 1890 
 — 7 spp. 
 
Iolaus   Hübner, [1819] 
  Sapphires 
 
 — 120 spp. 
Stugeta   Druce, 1891 
 — 7 spp. 
Etesiolaus   Stempffer & Bennett, 1959 
 — 3 spp. 
Pratapa   Moore, [1881] 
  Tufted Royals 
 
 — 6 spp. 
Tajuria   Moore, [1881] 
  Royals 
 
 — 32 spp. 
Dacalana   Moore, 1884 
 — 23 spp. 
Maneca   de Nicéville in Marshall & de Nicéville, 1890 
 — 1 sp. 
Creon   de Nicéville, [1896] 
 — 1 sp. 
Bullis   de Nicéville, 1897 
 — 3 spp. 
Sukidion   Druce, 1891 
 — 1 sp. 
Purlisa   Distant, 1881 
 — 1 sp. 
Jacoona   Distant, 1884 
 — 2 spp. 
Neocheritra   Distant, 1885 
 — 5 spp. 
Manto   de Nicéville, 1895 
 — 1 sp. 
Mantoides   Druce, 1896 
 — 1 sp. 
Thrix   Doherty, 1891 
 — 1 sp. 
Charana   de Nicéville in Marshall & de Nicéville, 1890 
  Mandarin Blues 
 
 — 2 spp. 
Suasa   de Nicéville in Marshall & de Nicéville, 1890 
 — 1 sp. 
 
Paruparo   Takanami, 1982 
 — 8 spp. 
Matsutaroa   Hayashi, Schröder & Treadaway, 1984 
 — 1 sp. 
Rachana   Eliot in Corbet & Pendlebury, 1978 
 — 5 spp. 
 
Semanga   Distant, 1884 
 — 2 spp. 
Mota   de Nicéville in Marshall & de Nicéville, 1890 
 — 1 sp. 
 
Zinaspa   de Nicéville in Marshall & de Nicéville, 1890 
 — 4 spp. 
Surendra   Moore, 1878 
  Acacia Blues 
 
 — 4 spp. 
 
Tribe Zesiini  
Zesiusinae  Swinhoe, 1911; 153; TG : Zesius   Hübner
 
 
 
 
Zesius   Hübner, [1819] 
 — 2 spp. 
 
Tribe Hypotheclini   Eliot, 1973 
 
 
Cheritra   Moore, [1881] 
  Imperials 
 
 — 4 spp. 
Ritra   de Nicéville in Marshall & de Nicéville, 1890 
 — 1 sp. 
Cheritrella   de Nicéville, 1887 
  Imperials 
 
 — 1 sp. 
Ticherra   de Nicéville, 1887 
 — 1 sp. 
Drupadia   Moore, 1884 
 — 12 spp. 
Ahmetia   Özdikmen, 2008 
 — 1 sp. 
 
Tribe Remelanini   Eliot, 1973 
 
 
Tribe Eumaeini   Doubleday, 1847 
 
 
Section Callophrys  
Section  Callophrys  (Eumaeini); [NL4A] , 125
 
  
 
Section Brangas  
Section  Brangas  (Eumaeini); [NL4A] , 118
 
  
 
Brangas   Hübner, [1819] 
 — 16 spp. 
Dabreras   Bálint in Bálint & Faynel, 2008 
 — 1 sp. 
Thaeides   Johnson, Kruse & Kroenlein, 1997 
 — 5 spp. 
Enos   Johnson, Kruse & Kroenlein, 1997 
 — 5 spp. 
Lamasina   Robbins, 2002 
 — 6 spp. 
Evenus   Hübner, [1819] 
 — 13 spp. 
 
Section Atlides  
Section  Atlides  (Eumaeini); [NL4A] , 119
 
  
 
Johnsonita   Salazar & Constantino, 1995 
 — 7 spp. 
Brevianta   Johnson, Kruse & Kroenlein, 1997 
 — 12 spp. 
Ianusanta   Bálint, 2011 
 — 1 sp. 
Micandra   Schatz, 1888 
 — 10 spp. 
Timaeta   Johnson, Kruse & Kroenlein, 1997 
 — 16 spp. 
Rhamma   Johnson, 1992 
 — 28 spp. 
Phothecla   Robbins in Lamas, 2004 
 — 3 spp. 
Salazaria   D'Abrera & Bálint, 2001 
 — 2 spp. 
Temecla   Robbins, 2004 
 — 6 spp. 
Ipidecla   Dyar, 1916 
 — 3 spp. 
Penaincisalia   Johnson, 1990 
 — 45 spp. 
Podanotum   Torres, Hall, Willmott & Johnson, 1996 
 — 10 spp. 
Lathecla   Robbins, 2004 
 — 7 spp. 
Busbiina   Robbins, 2004 
 — 1 sp. 
 
Section Thereus  
Section  Thereus  (Eumaeini); [NL4A] , 122
 
  
 
Section Satyrium  
Section  Satyrium  (Eumaeini); [NL4A] , 124
 
  
 
Section Thestius  
Section  Thestius  (Eumaeini); [NL4A] , 126
 
  
 
Section Allosmaitia  
Section  Allosmaitia  (Eumaeini); [NL4A] , 127
 
  
 
Section Strymon  
Section  Strymon  (Eumaeini); [NL4A] , 130
 
  
 
Tmolus   Hübner, [1819] 
 — 6 spp. 
Nicolaea   Johnson, 1993 
 — 26 spp. 
Ministrymon   Clench, 1961 
 — 20 spp. 
Exorbaetta   Johnson, Austin, Le Crom & Salazar, 1997 
 — 1 sp. 
Gargina   Robbins, 2004 
 — 8 spp. 
Siderus   Kaye, 1904 
 — 8 spp. 
Theclopsis   Godman & Salvin, [1887] 
 — 8 spp. 
Ostrinotes   Johnson, Austin, Le Crom & Salazar, 1997 
 — 9 spp. 
Strephonota   Johnson, Austin, Le Crom & Salazar, 1997 
 — 25 spp. 
 
Section Panthiades  
Section  Panthiades  (Eumaeini); [NL4A] , 134
 
  
 
Hypostrymon   Clench, 1961 
 — 4 spp. 
Marachina   Robbins, 2004 
 — 3 spp. 
Apuecla   Robbins, 2004 
 — 3 spp. 
Nesiostrymon   Clench, 1964 
 — 8 spp. 
Balintus   D'Abrera, 2001 
 — 1 sp. 
Aubergina   Johnson, 1991 
 — 5 spp. 
Terenthina   Robbins, 2004 
 — 2 spp. 
Iaspis   Kaye, 1904 
 — 10 spp. 
Celmia   Johnson, 1991 
 — 6 spp. 
Dicya   Johnson, 1991 
 — 6 spp. 
Trichonis   Hewitson, 1865 
 — 2 spp. 
 
Section Erora  
Section  Erora  (Eumaeini); [NL4A] , 136
 
  
 
Subtribe Calycopidina   Duarte & Robbins, 2010 
Section  Lamprospilus  (Eumaeini); [NL4A] , 127
 
  
 
Lamprospilus   Geyer, 1832 
 — 10 spp. 
Badecla   Duarte & Robbins, 2010 
 — 6 spp. 
Arzecla   Duarte & Robbins, 2010 
 — 10 spp. 
Arumecla   Robbins & Duarte in Lamas, 2004 
 — 4 spp. 
Camissecla   Robbins & Duarte in Lamas, 2004 
 — 11 spp. 
Electrostrymon   Clench, 1961 
 — 11 spp. 
Rubroserrata   Johnson & Kroenlein, 1993 
 — 2 spp. 
Ziegleria   Johnson, 1993 
 — 4 spp. 
Kisutam   Johnson & Kroenlein, 1993 
 — 2 spp. 
Calycopis   Scudder, 1876 
 — 65 spp. 
 
Tribe Catapaecilmatini   Eliot, 1973 
Catapaecilmini ; [MRS] , 638
 
  
 
Tribe Tomarini   Eliot, 1973 
 
 
Tribe Aphnaeini  
Section  Aphnaeus  (Aphnaeini); Eliot, 1973, Bull. Br. Mus. nat. Hist.  (Ent.) 28  (6) : 436 
Section  Pseudaletis  (Aphnaeini); Eliot, 1973, Bull. Br. Mus. nat. Hist.  (Ent.) 28  (6) : 436 
Aphnaeinae  (Lycaenidae); Korb & Bolshakov, 2011,  Eversmannia Suppl.  2  : 68; Boyle, Kaliszewska, Espeland, Suderman, Fleming, Heath & Pierce, 2015, Syst. Ent.  40  : 177
Aphnaeini  (Theclinae); Eliot, 1973, Bull. Br. Mus. nat. Hist.  (Ent.) 28  (6) : 381 , 435 ; Heath, 1997, Metamorphosis Occ. Suppl.  2  : 9; Vane-Wright & de Jong, 2003,  Zool. Verh. Leiden  343  : 130;  [AfrL ]
 
  
 
Phasis   Hübner, [1819] 
 — 4 spp. 
Tylopaedia   Tite & Dickson, 1973 
 — 1 sp. 
Argyraspodes   Tite & Dickson, 1973 
 — 1 sp. 
Aloeides   Hübner, [1819] 
 — 58 spp. 
Erikssonia   Trimen, 1891 
 — 4 spp. 
Trimenia   Tite & Dickson, 1973 
 — 5 spp. 
Aphnaeus   Hübner, [1819] 
  Silver Spots 
 
 — 35 spp. 
Axiocerses   Hübner, [1819] 
  Scarlets 
 
 — 19 spp. 
Zeritis   Boisduval, [1836] 
 — 6 spp. 
Crudaria   Wallengren, 1875 
 — 3 spp. 
Cigaritis   Donzel, 1847 
  Silverlines 
  Leopard Butterflies 
  Barred Blues 
 
 — 72 spp. 
Chloroselas   Butler, [1886] 
  Gems 
 
 — 13 spp. 
Lipaphnaeus   Aurivillius, 1916 
 — 4 spp. 
Chrysoritis   Butler, [1898] 
 — 42 spp. 
Vansomerenia   Heath, 1997 
 — 1 sp. 
Cesa   Seven, 1997 
 — 1 sp. 
Pseudaletis   Druce, 1888 
 — 29 spp. 
 
 
  
363x500 (~25Kb) Venezuela, Merida and the Sierra Nevada National Park, July 2002, Photo © Simon Coombes 
 
 
The exact identification of this species is still unknown,
but tentatively assumed to belong into this group. 
 Unknown or unplaced taxa
 
7.12.2023 (2654)
References: 
[AfrL] Mark C. Williams 
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007);  Afrototropical Butterflies ; e-mail:  Mark C. Williams 
 [CoL] Catalogue of Life 
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26);  https://doi.org/10.48580/dfz8d 
 [EVERSMANNIA]  
 Eversmannia contents 
 [NHM-BMW] Natural History Museum 
Butterflies and Moths of the World;  Generic Names and their Type-species 
 [Simon Coombes]  
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail:  Simon Coombes ;  http://www.butterfly-guide.co.uk 
 [UWSP] University of Wisconsin, Stevens Point 
 Reports of UWSP Museum of Natural History 
 [iNaturalist]  
 iNaturalist 
   
 
Some related literature: 
Boyle, Kaliszewska, Espeland, Suderman, Fleming, Heath & Pierce, 2015 
Phylogeny of the Aphnaeinae: myrmecophilous African butterflies with carnivorous and herbivorous life histories
Syst. Ent.  40 : 169-182
 [MRS]; Chou, 1994 
Monografia Rhopalocerorum Sinensium 1-2
Mon. Rhop. Sinesium 
: 1-854
 Duarte & Robbins, 2010 
Description and phylogenetic analysis of the Calycopidina (Lepidoptera, Lycaenidae, Theclinae, Eumaeini): a subtribe of detritivores
Revta Bras. Ent.  54  (1) 
: 45-65
 Eliot, 1973 
The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement
Bull. Br. Mus. nat. Hist.  (Ent.) 28  (6)
: 373 -505 (162fgs),  pl. 1 -6
 Clench in Fox, Lindsey, Clench & Miller, 1965 
The Butterflies of Liberia
Mem. Amer. Ent. Soc.  19 
: 1 -438
 Heath, 1997 
A review of African genera of the tribe Aphnaeini   (Lepidoptera: Lycaenidae)
Metamorphosis Occ. Suppl.  2 
: 1-60
 Johnson, 1993 
A New Neotropical Genus of Branded Angulopine-like Hairstreak Butterflies (Lepidoptera, Lycaenidae, Theclinae)
Rep. Mus. Nat. Hist. Univ. Wisc.  (Stevens Point) 40 
: 1-7
 Korb & Bolshakov, 2011 
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian)
Eversmannia Suppl.  2  
: 1-123
 [NL4A]; Lamas, 2004 
Checklist: Part 4A, Hesperioidea - Papilionoidea
Atlas of Neotropical Lepidoptera 
 [RFEL]; Leley, 2016 
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera
Cat. ins. Russian Far East.  2  
: 1-812
 Robbins & Busby, 2015 
Evolutionary gain of male secondary sexual structures in the widespread Neotropical montane genus Lathecla   (Lepidoptera, Lycaenidae, Eumaeini)
Insect Syst. Evol.  46 
: 47-78
 Strand, 1916 
Abbildungen und Beschreibungen neuer und wenig bekannter Lepidoptera aus der Sammlung W. Niepelt
Lepidoptera Niepeltiana  (2)
: 1 -26,  pl. 13 -17
 Vane-Wright & de Jong, 2003 
The butterflies of Sulawesi: annotated checklist for a critical island faunda
Zool. Verh. Leiden  343  : 3-268, pl. 1-16
   
 
If you have corrections, comments or
 information to add into these pages, just send mail to
Markku Savela  
Keep in mind that the taxonomic information is copied from various sources,  and may include many inaccuracies. Expert help is welcome.